The short-legged Andean cosmetids revisited: the genus Libitia Simon, 1879 with description of two new species (Opiliones, Cosmetidae)
Author
Medrano, Miguel
C7F7D4CF-F9B2-44AF-9F03-86278ADBD4F2
miangelmed@gmail.com
Author
Ázara, Ludson Neves de
4ECF193A-694C-43CE-8EE6-F197EDDA4414
Author
Kury, Adriano Brilhante
60FAE1F8-87F7-4A5F-BE78-BEB25BC4F898
text
European Journal of Taxonomy
2020
2020-04-24
634
1
25
journal article
22444
10.5852/ejt.2020.634
6531d570-f61a-49ae-a7a2-8167a41981fc
3778764
66AEE1D7-51BF-4583-9A19-947F61ECC7DE
Genus
Libitia
Simon, 1879
Libitia
Simon, 1879: 216
[
type
species:
Cosmetus cordatus
Gervais, 1844
, by subsequent designation of
Roewer (1912)
].
Libitiella
Roewer, 1947: 8
[
type
species:
Libitia bipunctata
Sørensen, 1932
, by original designation].
syn. nov.
Libitia
–
Simon 1880: 103
—
Roewer 1912: 11
;
1923: 293
. —
Mello-Leitão 1923: 108
;
1926: 331
;
1932: 53
;
1933: 105
, 109; 1935: 113. —
Kästner 1937: 389
. —
Kury 2003: 65
(revalidated) [junior subjective synonym of
Vonones
Simon, 1879
by
Goodnight & Goodnight (1953: 59)
].
Libitia
(
Libitia
)
– Sørensen
in
Henriksen 1932: 417.
non Libita
[subsequent incorrect spelling] –
Roewer 1928: 552
.
Etymology
Obscure, it sounds to us as a Roman woman’s name. Gender feminine.
Diagnosis
Can be differentiated in having shorter legs (femur IV shorter than the total length of the dorsal scutum) contrasting with other short-legged cosmetids as
Rhaucus
Simon, 1879
,
Neocynorta
or
Eulibitia
(its femur IV is longer than dorsal scutum length) (
Fig. 1B
, E–F). Outline of dorsal scutum alpha-type with faint marked constrictions forming a parallel coda in males and divergent coda in females contrasting with
Ambatoiella
that has well-marked constrictions (
Figs 2
,
6
). Can be separated from other genera by having cheliceral sockets (invaginations in the anterior margin of the carapace where the basichelicerites fit) shallow, with well-marked lateral quadrangular projections (triangular in
Eulibitia
, rounded in
Rhaucus
, bifid in
Metalibitia
) (
Figs 2
,
6
). Posterior margin of DS without yellow stripe (transversal stripe in
Eulibitia
and some
Rhaucus
species) (
Fig. 1
A–D). Basitarsomeres I, III and IV remarkably enlarged in males (
Figs 1B
,
5B, D
,
9
C–D). Penis with an elongate wattle in stylus forming distally two long serrate membranes and with a medial cleft (
Fig. 5
), without stylar barbs.
Description
Small and flat animals (<
4 mm
) with short legs (
Fig. 1B
). Outline of dorsal scutum alpha-type with symmetrical mid-bulge (
Fig. 2
).
Mesotergum flat and unarmed. Scutum with yellowish white blots as a ‘ladder mask’ over dark brown background (see
Medrano & Kury 2016
,
2017
), with intraspecific variation from absent (
Fig. 2
) to strongly filled invading the entire area I and II (
Fig. 6
) or part of carapace (
Fig. 9A
).
Coxa IV entirely visible in dorsal view, without groin warts (defined in
Kury & Barros, 2014
) and small prodorsal distal apophysis. Femur III and IV curved and uniformly tuberculate (
Fig. 1B
).
Penis (
Fig. 4
): ventral plate (VP) sub-rectangular with latero-apical borders protruding from the ventral plate, stylus with an incrassated wattle forming apically two long serrate membranes creating a medial cleft (
Fig. 4D
), without stylar barbs. VP laterally with two pairs of MS A, two pairs of MS C (sometimes asymmetrical with 3 MS C on one side, as in
L. bipunctata
and
L. gandalf
sp. nov.
), two pairs of MS D (D1 three times longer than D2), and ventrally with two pairs of MS E and one pair MS B (
Fig. 4C
).
Note
As no diagnostic differences have been found among the penes of each of the four species, no specific descriptions are made.
Included species
Libitia cordata
(
Gervais, 1844
)
,
Libitia bipunctata
Sørensen, 1932
,
Libitia gandalf
sp. nov.
and
Libitia iguaque
sp. nov.
Geographic distribution
Central part of Eastern Cordillera of Andes from 2600 to 3700 meters, in
Cundinamarca
,
Boyacá
and
Meta
departments of
Colombia
.