New species of Prototilla Schuster, 1949 from Argentina and diagnoses of the genus based on male and female (Hymenoptera: Bradynobaenidae: Typhoctinae: Eotillini) Author Torréns, Javier Author Fidalgo, Patricio Author Roig-Alsina, Arturo Author Brothers, Denis J. text Zootaxa 2012 3427 17 32 journal article 10.5281/zenodo.215056 9518f767-ef4a-4879-a21d-72b3f5f8233a 1175-5326 215056 Prototilla nigra Torréns, Fidalgo, Roig-Alsina & Brothers , sp. nov. ( Figs 10–16 ) Diagnosis. Male. General body color black ( Figs 11, 13 ). Scale-like setae covering posterior half of T1 and center of T2 ( Fig. 12 ). Fore wing barely setose ( Fig. 10 ). Female. Unknown. Description. Male. Length 5.0– 6.7 mm . General body color black. Scape, pedicel, trochanters and mandible dark brown; rest of legs and antennae light brown; last 4 segments of flagellum lighter than the rest ( Figs 11, 13, 14, 15 ). Head oval, 1.3–1.4 × as broad as high and 1.1–1.2 × as broad as mesosoma, with numerous scale-like setae covering almost entire head, vertex without or with a few scattered scale-like setae; plumose setae long, white and scattered over vertex and frons almost reaching toruli; vertex with spaced punctation ( Fig. 14 ). POL 1.0–1.1 × as long as OOL, POL 1.5–1.9 × as long as LOL; eyes ovoid, separated by 1.2–1.3 × their height, malar space 0.1–0.2 × eye height. Mandible arched, simple, with a small preapical denticle on inner edge; palpal formula 6- 4. Antenna of 13 segments; radicle as long as broad; scape with scale-like and plumose scattered setae, except ventrally glabrous, scale-like setae denser on external surface; scape 1.8–2.2 × as long as broad and 2.2–2.6 × as long as pedicel; flagellomeres cylindrical, F1 0.7–0.8 × length of scape and 0.8–0.9 × length of F2, F2–F6 approximately same length, F7–F10 gradually decreasing in length, F11 elongated with rounded apex ( Fig. 15 ). FIGURES 11–16. Prototilla nigra sp. nov. , holotype 3 (Lavalle, Mendoza): 11, habitus (lateral); 12, metasoma (dorsal); 13, habitus (dorsal); 14, head (anterior); 15, antenna (lateral); 16, genitalia (ventral; genital foramen projection indicated). Mesosoma elongate, 1.4–1.7 × as long as broad; punctation well marked, similar to that of head, spaced on dorsum and more contiguous on pleura and propodeum which becomes rugose toward insertion of metasoma; covered with scale-like and plumose setae ( Figs 11, 13 ). Pronotum transverse, anteriorly forming a collar with small plumose setae, rest with scale-like and white plumose setae, lateral surface with scattered setae and spaced punctation; humeral angle well marked but not acute. Mesonotum moderately punctate, with scattered scale-like and white plumose setae; tegula convex, smooth and with a few scale-like setae dorsally; scutellum with wellscattered punctation and with scale-like and plumose setae, except medially smooth and without setae. Metanotum transverse, narrow, with scale-like and plumose setae; punctate except medially smooth and without setae. Propodeum in dorsal view with spaced punctation and moderately dense scale-like setae, punctation becoming contiguous, deeper and wider on vertical surfaces, and with predominance of plumose setae; a group of scale-like setae at insertion of hind coxa. Propleuron heavily punctate, covered almost completely with scale-like setae; mesopleuron convex, punctate, densely covered by scale-like and plumose setae; metapleuron with punctation contiguous, covered with scale-like setae. Wing venation as for genus. Fore wing 3.3 × as long as broad, barely setose ( Fig. 10 ), hyaline, venation light brown, with white and thick plumose setae at base, becoming light brown on veins C and M+Cu . Legs slender; anterior surface of coxae, trochanters, femora and tibiae almost covered by scale-like setae, rest of their surfaces with or without scattered scale-like setae; plumose setae scattered across surfaces of legs, becoming denser where no scale-like setae; mid and hind tibiae each with two long white spurs, claws toothed. Metasoma 2.6–2.7 × as long as broad; terga with well-marked punctation slightly spaced on T1 and T2, remaining terga with punctation less marked and contiguous ( Fig. 12 ). Posterior margins of T1–T5 with scale-like setae; T1 covered with plumose setae, except covered by scale-like setae on posterior half; T2 with plumose setae covering all surfaces, denser laterally and scattered on dorsal surface at median group of scale-like setae; rest of terga with scattered plumose setae; posterior margin of T6 with dark brown, thick and apically truncate setae; T7 with surface convex; sterna with sculpture and distribution of setae similar to corresponding terga. Genitalia with paramere slender and curved upwards, with long setae along surface and small setae towards apex; volsella well developed, digitus swollen towards apex and cuspis thin, both with few long setae on surface and very small setae distally; aedeagus bipartite; basal ring anteroventrally narrowly and acutely emarginate ( Fig. 16 ). Female. Unknown. Variation. The antennae and legs are sometimes darker, and the femora may be almost completely dark brown; the predominance of scale-like setae on the mesosoma is higher in some specimens, especially on the metapleuron; on T2 the scale-like setae may cover almost the entire dorsal region. Etymology. From Latin niger -gra -grum which means “black”, referring to the general body color that is mostly black to dark brown. Distribution. Argentina : La Rioja, Mendoza. Material examined. Holotype 3: ARGENTINA : Mendoza, Lavalle, Reserva Telteca [ 32°23'6"S 68°3'0"W ], 15/XII/1994 – 03/II/1995 , G. Flores and S. Roig (IADIZA). Paratypes , 23: La Rioja, Santa Teresita [ 28°35'57"S 66°33'32"W ], XI/2005 , J. Torréns, G. Fidalgo and P. Fidalgo, TM (13, IFML ; 13, MACN ).