Characterization of Pleurotinae, with review of Pleurota species close to P. aristella (Linnaeus) from Morocco (Lepidoptera: Gelechioidea: Oecophoridae)
Author
Kaila, Lauri
Author
Mutanen, Marko
Author
Sihvonen, Pasi
Author
Tyllinen, Juha
Author
Tabell, Jukka
text
Zootaxa
2019
2019-01-21
4545
4
451
477
journal article
27677
10.11646/zootaxa.4545.4.1
36219e98-ffc6-46c4-86b1-41e0ff4c2215
1175-5326
2618231
107B7CF9-8A4E-4D27-86A3-02A0212285DF
Pleurota staintoniella
Baker, 1888
Barcode Index Number: BOLD:ADI3642
Table 1,
Figs. 20–21
,
30–31
,
42
,
47
,
48
Material studied.
12 ♂
(GP 5700 J. Tabell, DNA sample 25570 Lepid Phyl; GP 5730 J. Tabell; DNA sample 25571 Lepid Phyl),
4 ♀
(GP 5710
J. Tabell
, DNA sample 25569
Lepid Phyl
; GP 5899
J. Tabell
, DNA sample 25572
Lepid Phyl
):
Morocco
,
Taourirt Prov.
,
3 km
NNE Debdou
,
880 m
,
N34.00987
W3.02433
,
11.V.2016
,
J. Tabell
leg
. (coll.
TAB
).
The
male
lectotype
(
Algeria
, Sebdou) is deposited in NHMUK
.
Diagnosis.
Characterized by a unicolorous brown or yellowish brown forewing with white costal and median lines. In the male genitalia, a narrow and long phallus, and a long, well-sclerotized valval lobe of the juxta are the distinguishing features. In the female genitalia, extremely long apophyses and broad projections of the posterior signum are characteristic.
Molecular diagnosis.
Four specimens of
P. staintoniella
were sequenced, all of which yielded 658 bp DNA barcode fragments. The nearest neighbour to
P. staintoniella
is an undetermined Moroccan species (females only available) with a 5.09% divergence. The barcodes of
P. staintoniella
exhibit 0.46 maximum intraspecific variation (Table 1).
FIGURE 42.
Male genitalia.
P.
staintoniella
Baker
, GP 5700 J. Tabell.
Description.
Adult. Wingspan
19–25 mm
. Labial palpus off-white, mixed brown, long, about 6.7 x as long as diameter of eye (1
st
and 2
nd
segments), 3
rd
segment long, 0.38 x length of 1
st
and 2
nd
segments. Antenna brown. Head whitish tinged pale brown, tegula and thorax concolorous with forewing. Forewing pale yellowish brown; costal line white, narrow, from near apex to 0.8; median line white, extending from base to apex, slightly expanded medially, dorso- medially with two shallow notches, apical 0.4 narrow. Fringe brown mixed white. Hindwing brownish grey, margin brown, fringe brown, apically paler. Abdomen slightly lustrous, greyish brown, each segment with transverse row of ochre scales.
FIGURES 43–45.
Female genitalia. 43.
P. lacteella
sp. nov.
, GP 5429 J. Tabell. 44.
P. moroccoensis
sp. nov.
, GP 5456 J. Tabell. 45.
P. atlasensis
sp. nov.
, GP 5545 J. Tabell.
FIGURES 46–47.
Female genitalia. 46.
P. ochreopalpella
sp. nov.
, GP 5898 J. Tabell. 47
P. staintoniella
Baker
, ♀ genitalia, GP 5899 J. Tabell.
FIGURE 48.
Neighbour joining tree, conducted under Kimura 2-parameter model, of ten taxa of
Pleurota aristella
group, based on sequences of
COI
gene. The values by the nodes indicate node bootstrap supports as based on 500 replicates.
Male genitalia. Uncus thimble-shaped dorsoventrally, lined with several long bristles, apical 0.5 more strongly sclerotized. Gnathos long, almost
2 x
as long as uncus, funnel-shaped dorsoventrally, beak-shaped in lateral aspect, medially covered with small spines. Valva upwards oblique, ventral margin slightly convex, costa straight; sacculus narrow and long. Anterior extension of juxta straight; median part chaliced; posterior lobe very long, exceeding apex of uncus; valval lobe long, club-shaped, smooth, distally covered with several small nodules and short bristles. Vinculum U-shaped. Phallus curved, parallel-sided, apex blunt, with one plate-shaped cornutus, apical 0.14 dorsally covered with small nodules.
Female genitalia. Papillae anales ovoid, covered with bristles. Apophyses very long; posterior apophysis 5.5 x as long as papilla analis and 1.7 x as long as anterior apophysis. Segment 8 elongated longitudinally, rectangular, membranous, dorsolaterally reinforced by sclerotized band, ventrally sparsely covered with bristles, ventral longitudinal sclerotization narrow, slightly shorter than segment 8, branch of apophysis anterioris narrow and short. Antrum tubular, parallel-sided, shorter than segment 8, posteriorly membranous, anterior 0.33 tapered and sclerotized. Ductus bursae membranous, as long as antrum, slightly expanded towards corpus bursae. Corpus bursae ovoid with one robust arched signum with two long, narrow, sclerotized thorn-like protuberances, and two elongate leaf-shaped signa.
Biology.
Unknown. In
Morocco
, the specimens were collected on a calcareous hill in May, together with
P. ochreopalpella
.
Distribution.
Previously only known from
Algeria
. The distance between the Algerian
type
locality and Moroccan site is less than
200 km
.