Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae) Author Reynoso-Velasco, Daniel Author Sites, Robert W. text Zootaxa 2019 2019-06-13 4615 2 285 302 journal article 26498 10.11646/zootaxa.4615.2.3 a94be2b4-e3c7-4a35-a5e8-5776197d8ca6 1175-5326 3244686 2222331C-04B8-4C64-B190-C8AE3A57D44E Ambrysus signoreti Stål species complex Usinger (1946) first made reference to the “ signoreti ” complex when he described 16 new species in Ambrysus . In a later work, La Rivers (1957) listed the unifying features of species in the A. signoreti complex: 1) broad body shape, 2) wide embolia, 3) prominent maculation, and 4) posterolateral corners of abdominal segments III–V produced (spinose).