<document id="A56BD21A55D6A7EEA39B006539A2D32C" ID-CLB-Dataset="20271" ID-DOI="10.5852/ejt.2022.799.1681" ID-GBIF-Dataset="f53d5465-7162-45d6-892b-dfc0b8d99789" ID-ISSN="2118-9773" ID-Zenodo-Dep="6341270" ID-ZooBank="F52DBAB0-38C7-400F-9BA1-E2D8E6B19E7E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1646831675572" checkinUser="felipe" docAuthor="Aubrechtová, Martina & Korn, Dieter" docDate="2022" docId="03FE5868FFB63D1FDD0EFCE1FE32F9D1" docLanguage="en" docName="ejt-799_1-108.pdf" docOrigin="European Journal of Taxonomy 799 (1)" docSource="http://dx.doi.org/10.5852/ejt.2022.799.1681" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Lituites fallax Remele 1890" docType="treatment" docVersion="10" lastPageNumber="69" masterDocId="FFC72010FFF53D5BDF7EFF83FFE3FFD1" masterDocTitle="Taxonomy and ontogeny of the Lituitida (Cephalopoda) from Orthoceratite Limestone erratics (Middle Ordovician)" masterLastPageNumber="108" masterPageNumber="1" pageNumber="68" updateTime="1699317703155" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED"> <mods:mods id="2EB50710AA7DDABB53D080BB58E139B0" xmlns:mods="http://www.loc.gov/mods/v3"> <mods:titleInfo id="7070D2153AD3705BC2F578836F101973"> <mods:title id="55EC21B6352361992F39C3B08BC9B3B9">Taxonomy and ontogeny of the Lituitida (Cephalopoda) from Orthoceratite Limestone erratics (Middle Ordovician)</mods:title> </mods:titleInfo> <mods:name id="BBED908D2B292FD0CC72649DBA369ABD" type="personal"> <mods:role id="A03FD9E8DD83CFF526F47F0366E4B88F"> <mods:roleTerm id="CCF6597BD7F179A3954D434332CFF534">Author</mods:roleTerm> </mods:role> <mods:namePart id="80404FFCDC45187007E8076CFC93770E">Aubrechtová, Martina</mods:namePart> <mods:nameIdentifier id="F01F47B0420AC153881740D255020FD4" type="LSID">CAF4231-8787-4051-8D76-F983332517EE</mods:nameIdentifier> <mods:affiliation id="6F807D5C613A58FCC210EE19487CA7AC">Institute of Geology and Palaeontology, Faculty of Science, Charles University Prague, Albertov 6, Prague, 12843, Czech Republic. & Institute of Geology, Czech Academy of Sciences, Rozvojová 269, Prague, 16500, Czech Republic.</mods:affiliation> <mods:nameIdentifier id="687BA31CD6938B3A07C946F562D45F26" type="email">aubrech1@natur.cuni.cz,aubrechtova@gli.cas.cz</mods:nameIdentifier> </mods:name> <mods:name id="55BAAC0DF05B4EF054DB341F179DB02D" type="personal"> <mods:role id="634190DCDAA87536FDB92732451F0D30"> <mods:roleTerm id="652F268F32812D89DEFA676D5C635D0D">Author</mods:roleTerm> </mods:role> <mods:namePart id="6119322B7F60FC4A1378CD92D5083996">Korn, Dieter</mods:namePart> <mods:nameIdentifier id="FD25DD0FB2F05BDF72893661F7CA059C" type="LSID">286CA4F3-7EBC-4AEF-A66A-B2508D001367</mods:nameIdentifier> <mods:affiliation id="C6088F183715242D598FA85CB701C6F1">Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation> <mods:nameIdentifier id="6381FCEF4294DF548EA346E840752FCA" type="email">dieter.korn@mfn.berlin</mods:nameIdentifier> </mods:name> <mods:typeOfResource id="8102EA1924E865CF5C22B1E130476A59">text</mods:typeOfResource> <mods:relatedItem id="6691B79FFA1577715F545E3BD0D908B0" type="host"> <mods:titleInfo id="FC225971BF9389D4E67AC95D9FEAF224"> <mods:title id="37EEB5D0139EB03A24F72627655F46CA">European Journal of Taxonomy</mods:title> </mods:titleInfo> <mods:part id="C682C368007653FDF66D47F18456F8D5"> <mods:date id="118EF3EF15736FF663EE1CECD3934ADB">2022</mods:date> <mods:detail id="A8B99A58B7EA2491A0ACFEA8365E7949" type="pubDate"> <mods:number id="75F005060647C56ABDC37CB463549497">2022-03-08</mods:number> </mods:detail> <mods:detail id="1500F58DDFE0F4AAE6FC0A97C2D3F909" type="volume"> <mods:number id="1C73F8DDA45AFB8594B327981302DFF1">799</mods:number> </mods:detail> <mods:detail id="23F5E3F905B65284C1452C108B923DDF" type="issue"> <mods:number id="DAC96281D84F378ADA91E53F8FBD8025">1</mods:number> </mods:detail> <mods:extent id="8F0515C1245E42B8DA938DE59926C396" unit="page"> <mods:start id="1C9B3C0FDCD826A1AC157F0BEA9CCD76">1</mods:start> <mods:end id="597C399487BC2B6FC2F529C5C5DF66F0">108</mods:end> </mods:extent> </mods:part> </mods:relatedItem> <mods:location id="BA42A4E2D4167FE7998F87DD53EDBA9A"> <mods:url id="1937AAE292572E82973B36A62B77FACE">http://dx.doi.org/10.5852/ejt.2022.799.1681</mods:url> </mods:location> <mods:classification id="3FAEB709C3C85FEABDF7DC6E657BE6E4">journal article</mods:classification> <mods:identifier id="0FF74DDB7E3C979D5FA34E4B7C35BC96" type="CLB-Dataset">20271</mods:identifier> <mods:identifier id="F46DF11A3136C29C056E293BC4D1C219" type="DOI">10.5852/ejt.2022.799.1681</mods:identifier> <mods:identifier id="8660B64B72364342A65F36D5309ADCBA" type="GBIF-Dataset">f53d5465-7162-45d6-892b-dfc0b8d99789</mods:identifier> <mods:identifier id="35DBF14FD5E4A0BF44110302268C96FC" type="ISSN">2118-9773</mods:identifier> <mods:identifier id="1992F647843F23D207097D73B28B908A" type="Zenodo-Dep">6341270</mods:identifier> <mods:identifier id="F46A7F02260D75BBFE455EDBFBA3918E" type="ZooBank">F52DBAB0-38C7-400F-9BA1-E2D8E6B19E7E</mods:identifier> </mods:mods> <treatment id="03FE5868FFB63D1FDD0EFCE1FE32F9D1" ID-DOI="http://doi.org/10.5281/zenodo.6343264" ID-GBIF-Taxon="193941446" ID-Zenodo-Dep="6343264" LSID="urn:lsid:plazi:treatment:03FE5868FFB63D1FDD0EFCE1FE32F9D1" httpUri="http://treatment.plazi.org/id/03FE5868FFB63D1FDD0EFCE1FE32F9D1" lastPageId="68" lastPageNumber="69" pageId="67" pageNumber="68"> <subSubSection id="C34DBAF5FFB63D18DD0EFCE1FC27FCAD" box="[624,964,866,893]" pageId="67" pageNumber="68" type="nomenclature"> <paragraph id="8BE8E97EFFB63D18DD0EFCE1FC27FCAD" blockId="67.[624,964,866,927]" box="[624,964,866,893]" pageId="67" pageNumber="68"> <heading id="D0A05E12FFB63D18DD0EFCE1FC27FCAD" box="[624,964,866,893]" centered="true" fontSize="11" level="2" pageId="67" pageNumber="68" reason="2"> <taxonomicName id="4C5792FDFFB63D18DD0EFCE1FC27FCAD" authority="Remele, 1890" authorityName="Remele" authorityYear="1890" box="[624,964,866,893]" class="Cephalopoda" family="Lituitidae" genus="Lituites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="67" pageNumber="68" phylum="Mollusca" rank="species" species="fallax"> <emphasis id="B923356CFFB63D18DD0EFCE1FCFAFCAC" bold="true" box="[624,793,866,893]" italics="true" pageId="67" pageNumber="68">Lituites fallax</emphasis> Remelé, 1890 </taxonomicName> </heading> </paragraph> </subSubSection> <subSubSection id="C34DBAF5FFB63D18DDC0FC06FC95FC4E" box="[702,886,901,927]" pageId="67" pageNumber="68" type="description"> <paragraph id="8BE8E97EFFB63D18DDC0FC06FC95FC4E" blockId="67.[624,964,866,927]" box="[702,886,901,927]" pageId="67" pageNumber="68"> <figureCitation id="136CF5FBFFB63D18DDC0FC06FCF7FC4E" box="[702,788,901,927]" captionStart="Fig" captionStartId="68.[189,232,950,976]" captionTargetBox="[295,1295,272,906]" captionTargetId="figure-260@68.[285,1303,265,913]" captionTargetPageId="68" captionText="Fig. 46. Lituites fallax Remelé, 1890 from the Upper Grey Orthoceratite Limestone. Lectotype MB.C.11672.2 (Ramann, Kgl. Forstakademie Eberswalde Coll.) from Heegermühle (Brandenburg, Germany); photograph (with dorsal, lateral and ventral views) and reproduction of the illustration by Remelé (1890: pl. 5 fig. 1). Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6341370" httpUri="https://zenodo.org/record/6341370/files/figure.png" pageId="67" pageNumber="68">Fig. 46</figureCitation> , <tableCitation id="C6D5DCC5FFB63D18DC61FC06FC95FC4E" box="[799,886,901,927]" captionStart="Table 9" captionStartId="52.[189,259,265,291]" captionTargetBox="[200,1358,367,1950]" captionTargetPageId="52" captionText="Table 9. Conch measurements (in mm), ratios and rates of coiled conch parts of Lituites Bertrand, 1763." pageId="67" pageNumber="68">Table 9</tableCitation> </paragraph> </subSubSection> <subSubSection id="C34DBAF5FFB63D18DFC3FC4BFD27FC30" box="[189,708,967,994]" pageId="67" pageNumber="68" type="reference_group"> <paragraph id="8BE8E97EFFB63D18DFC3FC4BFD27FC30" blockId="67.[189,708,967,994]" box="[189,708,967,994]" pageId="67" pageNumber="68"> <treatmentCitationGroup id="AB47CE50FFB63D18DFC3FC4BFD27FC30" box="[189,708,967,994]" pageId="67" pageNumber="68"> <treatmentCitation id="0AF6CF6FFFB63D18DFC3FC4BFD27FC30" author="Remelé" box="[189,708,967,994]" page="59" pageId="67" pageNumber="68" year="1890"> <taxonomicName id="4C5792FDFFB63D18DFC3FC4BFDDAFC33" authority="Remele, 1890: 59" authorityName="Remele" authorityPageNumber="59" authorityYear="1890" box="[189,569,967,994]" class="Cephalopoda" family="Lituitidae" genus="Lituites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="67" pageNumber="68" phylum="Mollusca" rank="species" species="fallax"> <emphasis id="B923356CFFB63D18DFC3FC4BFEBCFC30" box="[189,351,967,993]" italics="true" pageId="67" pageNumber="68">Lituites fallax</emphasis> Remelé, 1890: 59 </taxonomicName> , pl. 5 fig. 1. </treatmentCitation> </treatmentCitationGroup> </paragraph> </subSubSection> <subSubSection id="C34DBAF5FFB63D18DFC3FB8BFB4AFB46" pageId="67" pageNumber="68" type="diagnosis"> <paragraph id="8BE8E97EFFB63D18DFC3FB8BFED5FBF3" blockId="67.[189,310,1032,1058]" box="[189,310,1032,1058]" pageId="67" pageNumber="68"> <heading id="D0A05E12FFB63D18DFC3FB8BFED5FBF3" bold="true" box="[189,310,1032,1058]" fontSize="11" level="3" pageId="67" pageNumber="68" reason="3"> <emphasis id="B923356CFFB63D18DFC3FB8BFED5FBF3" bold="true" box="[189,310,1032,1058]" pageId="67" pageNumber="68">Diagnosis</emphasis> </heading> </paragraph> <paragraph id="8BE8E97EFFB63D18DFC3FBB4FB4AFB46" blockId="67.[189,1398,1079,1176]" pageId="67" pageNumber="68"> Species of the genus <taxonomicName id="4C5792FDFFB63D18DECAFBBBFDEFFB80" authorityName="Lasaulxii Remele" authorityYear="1890" box="[436,524,1080,1105]" class="Cephalopoda" family="Lituitidae" genus="Lituites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="67" pageNumber="68" phylum="Mollusca" rank="genus"> <emphasis id="B923356CFFB63D18DECAFBBBFDEFFB80" box="[436,524,1080,1105]" italics="true" pageId="67" pageNumber="68">Lituites</emphasis> </taxonomicName> with coiled conch diameter of ca <quantity id="4CAF449BFFB63D18DCE4FBB4FC0CFB80" box="[922,1007,1079,1105]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="67" pageNumber="68" unit="mm" value="40.0">40 mm</quantity> ; whorl expansion rate 1.70. Shell surface of the coiled part of lirae (0.5–1.0 mm apart), raised in ca <quantity id="4CAF449BFFB63D18DCB6FBD9FBEFFBA4" box="[968,1036,1114,1141]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="67" pageNumber="68" unit="mm" value="2.0">2 mm</quantity> wide intervals. Deep, rounded ventral sinus, very low dorsal projection; lirae straight laterally, direction rectiradiate. </paragraph> </subSubSection> <subSubSection id="C34DBAF5FFB63D18DFC3FB3CFCFCFAA0" pageId="67" pageNumber="68" type="materials_examined"> <paragraph id="8BE8E97EFFB63D18DFC3FB3CFE8FFB09" blockId="67.[189,364,1214,1241]" box="[189,364,1214,1241]" pageId="67" pageNumber="68"> <heading id="D0A05E12FFB63D18DFC3FB3CFE8FFB09" bold="true" box="[189,364,1214,1241]" fontSize="11" level="3" pageId="67" pageNumber="68" reason="3"> <emphasis id="B923356CFFB63D18DFC3FB3CFE8FFB09" bold="true" box="[189,364,1214,1241]" pageId="67" pageNumber="68">Type material</emphasis> </heading> </paragraph> <paragraph id="8BE8E97EFFB63D18DF94FB6EFCFCFAA0" blockId="67.[189,1399,1261,1393]" pageId="67" pageNumber="68"> <materialsCitation id="3B3FE323FFB63D18DF94FB6EFCFCFAA0" ID-GBIF-Occurrence="3700114361" collectionCode="MB" collectorName="Ramann" country="Germany" county="Heegermuhle" location="Heegermuhle" municipality="Ordovician" pageId="67" pageNumber="68" specimenCode="MB.C.11672.2" specimenCount="1" stateProvince="Brandenburg" typeStatus="lectotype"> <emphasis id="B923356CFFB63D18DF94FB6EFE85FAD6" bold="true" box="[234,358,1261,1287]" pageId="67" pageNumber="68"> <typeStatus id="54EC57DCFFB63D18DF94FB6EFE85FAD6" box="[234,358,1261,1287]" pageId="67" pageNumber="68" type="lectotype">Lectotype</typeStatus> </emphasis> (designated herein) <collectingCountry id="F340A9EEFFB63D18DFC3FA93FEB3FAFB" box="[189,336,1296,1322]" name="Germany" pageId="67" pageNumber="68">GERMANY</collectingCountry> • <collectingRegion id="4993279CFFB63D18DE1CFA93FE1DFAFB" box="[354,510,1296,1322]" country="Germany" name="Brandenburg" pageId="67" pageNumber="68">Brandenburg</collectingRegion> , <collectingCounty id="628991F2FFB63D18DD78FA93FD4BFAFB" box="[518,680,1296,1322]" pageId="67" pageNumber="68">Heegermühle</collectingCounty> ; <geologicalTimeScale id="F97769D5FFB63D18DDCFFA93FA91FAFA" box="[689,1394,1296,1323]" pageId="67" pageNumber="68">Ordovician (late Lasnamägi Regional Stage, late Darriwilian)</geologicalTimeScale> , <geologicalTimeScale id="F97769D5FFB63D18DFC3FAB7FD63FA9C" box="[189,640,1331,1358]" pageId="67" pageNumber="68">Upper Grey Orthoceratite Limestone</geologicalTimeScale> ; <collectorName id="26A28CA8FFB63D18DDEEFAB7FD11FA9F" box="[656,754,1332,1358]" pageId="67" pageNumber="68">Ramann</collectorName> Coll.; previously illustrated by Remelé (1890: pl. 5 fig. 1), re-illustrated here in <figureCitation id="136CF5FBFFB63D18DD79FAD4FDBDFAA0" box="[519,606,1367,1393]" captionStart="Fig" captionStartId="68.[189,232,950,976]" captionTargetBox="[295,1295,272,906]" captionTargetId="figure-260@68.[285,1303,265,913]" captionTargetPageId="68" captionText="Fig. 46. Lituites fallax Remelé, 1890 from the Upper Grey Orthoceratite Limestone. Lectotype MB.C.11672.2 (Ramann, Kgl. Forstakademie Eberswalde Coll.) from Heegermühle (Brandenburg, Germany); photograph (with dorsal, lateral and ventral views) and reproduction of the illustration by Remelé (1890: pl. 5 fig. 1). Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6341370" httpUri="https://zenodo.org/record/6341370/files/figure.png" pageId="67" pageNumber="68">Fig. 46</figureCitation> ; <specimenCode id="DBF14105FFB63D18DD14FAD4FCFFFAA0" box="[618,796,1367,1393]" pageId="67" pageNumber="68">MB.C.11672.2</specimenCode> . </materialsCitation> </paragraph> </subSubSection> <subSubSection id="C34DBAF5FFB63D18DFC3FA1BFE16F9BC" pageId="67" pageNumber="68" type="description"> <paragraph id="8BE8E97EFFB63D18DFC3FA1BFEADFA63" blockId="67.[189,334,1432,1458]" box="[189,334,1432,1458]" pageId="67" pageNumber="68"> <heading id="D0A05E12FFB63D18DFC3FA1BFEADFA63" bold="true" box="[189,334,1432,1458]" fontSize="11" level="3" pageId="67" pageNumber="68" reason="3"> <emphasis id="B923356CFFB63D18DFC3FA1BFEADFA63" bold="true" box="[189,334,1432,1458]" pageId="67" pageNumber="68">Description</emphasis> </heading> </paragraph> <paragraph id="8BE8E97EFFB63D18DFC3FA44FE16F9BC" blockId="67.[188,1399,1479,1645]" pageId="67" pageNumber="68"> <typeStatus id="54EC57DCFFB63D18DFC3FA44FED1FA30" box="[189,306,1479,1505]" pageId="67" pageNumber="68" type="lectotype">Lectotype</typeStatus> MB.C.11672.2 ( <figureCitation id="136CF5FBFFB63D18DE84FA44FDB2FA33" box="[506,593,1479,1506]" captionStart="Fig" captionStartId="68.[189,232,950,976]" captionTargetBox="[295,1295,272,906]" captionTargetId="figure-260@68.[285,1303,265,913]" captionTargetPageId="68" captionText="Fig. 46. Lituites fallax Remelé, 1890 from the Upper Grey Orthoceratite Limestone. Lectotype MB.C.11672.2 (Ramann, Kgl. Forstakademie Eberswalde Coll.) from Heegermühle (Brandenburg, Germany); photograph (with dorsal, lateral and ventral views) and reproduction of the illustration by Remelé (1890: pl. 5 fig. 1). Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6341370" httpUri="https://zenodo.org/record/6341370/files/figure.png" pageId="67" pageNumber="68">Fig. 46</figureCitation> ) is a fragment of a third of the last whorl of the coiled part ( <quantity id="4CAF449BFFB63D18DA5CFA44FA94FA30" box="[1314,1399,1479,1505]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="67" pageNumber="68" unit="mm" value="40.0">40 mm</quantity> estimated diameter; WER <subScript id="17D3EB3BFFB63D18DE94FA7EFE19F9DA" attach="left" box="[490,506,1533,1547]" fontSize="6" pageId="67" pageNumber="68">ah</subScript> ~ 1.70) of a conch. The ornament consists of sharp lirae (0.5–1.0 mm apart), raised in more or less regular intervals (every ca <quantity id="4CAF449BFFB63D18DD90F98EFCD5F9F9" box="[750,822,1549,1576]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="67" pageNumber="68" unit="mm" value="2.0">2 mm</quantity> ); the lirae are straight on the flanks, with a narrow ventrolateral projection. The ventral sinus is deep and rounded, dorsal projection is very low with a shallow dorsolateral sinus. </paragraph> </subSubSection> <subSubSection id="C34DBAF5FFB63D1FDFC3F916FE32F9D1" lastPageId="68" lastPageNumber="69" pageId="67" pageNumber="68" type="discussion"> <paragraph id="8BE8E97EFFB63D18DFC3F916FECCF97E" blockId="67.[189,303,1685,1711]" box="[189,303,1685,1711]" pageId="67" pageNumber="68"> <heading id="D0A05E12FFB63D18DFC3F916FECCF97E" bold="true" box="[189,303,1685,1711]" fontSize="11" level="3" pageId="67" pageNumber="68" reason="3"> <emphasis id="B923356CFFB63D18DFC3F916FECCF97E" bold="true" box="[189,303,1685,1711]" pageId="67" pageNumber="68">Remarks</emphasis> </heading> </paragraph> <paragraph id="8BE8E97EFFB63D1FDFC3F940FDECFB3D" blockId="67.[188,1401,1731,2005]" lastBlockId="68.[189,1399,1163,1260]" lastPageId="68" lastPageNumber="69" pageId="67" pageNumber="68"> The <typeStatus id="54EC57DCFFB63D18DF8AF940FE83F90C" box="[244,352,1731,1757]" pageId="67" pageNumber="68" type="lectotype">lectotype</typeStatus> of <taxonomicName id="4C5792FDFFB63D18DEF0F947FDD0F90C" authorityName="Remele" authorityYear="1890" box="[398,563,1731,1757]" class="Cephalopoda" family="Lituitidae" genus="Lituites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="67" pageNumber="68" phylum="Mollusca" rank="species" species="fallax"> <emphasis id="B923356CFFB63D18DEF0F947FDD0F90C" box="[398,563,1731,1757]" italics="true" pageId="67" pageNumber="68">Lituites fallax</emphasis> </taxonomicName> is incompletely preserved and therefore, it cannot be excluded that it represents a morphological variant of some other species of the genus <emphasis id="B923356CFFB63D18DC8AF964FBB7F8D1" box="[1012,1108,1767,1792]" italics="true" pageId="67" pageNumber="68"> <taxonomicName id="4C5792FDFFB63D18DC8AF964FBB3F8D1" authorityName="Lasaulxii Remele" authorityYear="1890" box="[1012,1104,1767,1792]" class="Cephalopoda" family="Lituitidae" genus="Lituites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="67" pageNumber="68" phylum="Mollusca" rank="genus">Lituites</taxonomicName> . </emphasis> Most similar to <taxonomicName id="4C5792FDFFB63D18DA6BF964FA94F8D0" authorityName="Remele" authorityYear="1890" box="[1301,1399,1767,1793]" class="Cephalopoda" family="Lituitidae" genus="Lituites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="67" pageNumber="68" phylum="Mollusca" rank="species" species="fallax"> <emphasis id="B923356CFFB63D18DA6BF964FA94F8D0" box="[1301,1399,1767,1793]" italics="true" pageId="67" pageNumber="68">L. fallax</emphasis> </taxonomicName> are <taxonomicName id="4C5792FDFFB63D18DF99F888FE8AF8F5" authority="Aubrechtová & Korn, 2022" authorityName="Aubrechtová & Korn" authorityYear="2022" box="[231,361,1802,1828]" class="Cephalopoda" family="Lituitidae" genus="Lituites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="67" pageNumber="68" phylum="Mollusca" rank="species" species="kruegeri" status="sp. nov."> <emphasis id="B923356CFFB63D18DF99F888FE8AF8F5" box="[231,361,1802,1828]" italics="true" pageId="67" pageNumber="68">L. kruegeri</emphasis> </taxonomicName> <taxonomicNameLabel id="A2108817FFB63D18DE0EF889FE29F8F5" box="[368,458,1802,1828]" pageId="67" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel> and <taxonomicName id="4C5792FDFFB63D18DD7EF888FD44F8F5" authorityName="Aubrechtová & Korn" authorityYear="2022" box="[512,679,1802,1828]" class="Cephalopoda" family="Lituitidae" genus="Lituites" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Orthocerida" pageId="67" pageNumber="68" phylum="Mollusca" rank="species" species="kruegeri"> <emphasis id="B923356CFFB63D18DD7EF888FDFBF8F5" box="[512,536,1803,1828]" italics="true" pageId="67" pageNumber="68">L.</emphasis> cf. <emphasis id="B923356CFFB63D18DD3AF889FD44F8F5" box="[580,679,1802,1828]" italics="true" pageId="67" pageNumber="68">kruegeri</emphasis> </taxonomicName> , both of which differ from <taxonomicName id="4C5792FDFFB63D18DC9DF888FBA6F8F5" authorityName="Remele" authorityYear="1890" box="[995,1093,1802,1828]" class="Cephalopoda" family="Lituitidae" genus="Lituites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="67" pageNumber="68" phylum="Mollusca" rank="species" species="fallax"> <emphasis id="B923356CFFB63D18DC9DF888FBA6F8F5" box="[995,1093,1802,1828]" italics="true" pageId="67" pageNumber="68">L. fallax</emphasis> </taxonomicName> in having a smaller conch diameter (33 and <quantity id="4CAF449BFFB63D18DEF4F8AEFE03F899" box="[394,480,1837,1864]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="67" pageNumber="68" unit="mm" value="28.0">28 mm</quantity> , respectively) and a higher WER <subScript id="17D3EB3BFFB63D18DC1AF8C3FC97F89F" attach="left" box="[868,884,1856,1870]" fontSize="6" pageId="67" pageNumber="68">ah</subScript> (2 and 2.30, respectively). Also, <taxonomicName id="4C5792FDFFB63D18DB84F8ADFABFF896" authorityName="Remele" authorityYear="1890" box="[1274,1372,1837,1863]" class="Cephalopoda" family="Lituitidae" genus="Lituites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="67" pageNumber="68" phylum="Mollusca" rank="species" species="fallax"> <emphasis id="B923356CFFB63D18DB84F8ADFABFF896" box="[1274,1372,1837,1863]" italics="true" pageId="67" pageNumber="68">L. fallax</emphasis> </taxonomicName> is ornamented with lirae, which are straight on the flanks, while prorsiradiate lirae and flat annuli are developed in <taxonomicName id="4C5792FDFFB63D18DE1EF8F7FE07F85C" authority="Aubrechtová & Korn, 2022" authorityName="Aubrechtová & Korn" authorityYear="2022" box="[352,484,1907,1933]" class="Cephalopoda" family="Lituitidae" genus="Lituites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="67" pageNumber="68" phylum="Mollusca" rank="species" species="kruegeri" status="sp. nov."> <emphasis id="B923356CFFB63D18DE1EF8F7FE07F85C" box="[352,484,1907,1933]" italics="true" pageId="67" pageNumber="68">L. kruegeri</emphasis> </taxonomicName> <taxonomicNameLabel id="A2108817FFB63D18DE93F8F7FDAAF85F" box="[493,585,1908,1934]" pageId="67" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel> ( <figureCitation id="136CF5FBFFB63D18DD25F8F0FD41F85F" box="[603,674,1907,1934]" captionStart="Fig" captionStartId="9.[187,230,1640,1666]" captionTargetBox="[187,1383,620,1594]" captionTargetId="figure-89@9.[187,1396,610,1603]" captionTargetPageId="9" captionText="Fig. 8. Ornament details of lituitid specimens. A. Coarse lirae, symmetric crests; Rhynchorthoceras angelini (Boll, 1857), specimen MB.C.30495.1. B. Coarse lirae, asymmetric crests; R. angelini (Boll, 1857), specimen MB.C.11716. C. Faint, widestanding striae; R. rugium sp. nov., holotype MB.C.11728. D. Lirae and almost smooth internal shell; Ancistroceras torelli (Remelé, 1881), specimen MB.C.30519. E. Lirae and undulated internal shell; A. undulatum (Boll, 1857), specimen MB.C.11697. F. Narrow annuli, raised lirae and growth lines; Cyclolituites applanatus (Remelé, 1880), specimen MB.C.30560. G. Wide-standing, prorsiradiate lirae and very fine growth lines; Lituites dewitzi sp. nov., holotype MB.C.30538. H. Sharp lirae, straight on the flanks; L. fallax Remelé, 1890, holotype MB.C.11672.2. I. Sharp, prorsiradiate lirae and flat annuli; L. kruegeri sp. nov., holotype MB.C.30539.J. Wide-standing, sharp lirae and very flat annuli, straight on the flanks; Trilacinoceras knoefleri sp. nov., paratype MB.C.30559. Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6341289" httpUri="https://zenodo.org/record/6341289/files/figure.png" pageId="67" pageNumber="68">Fig. 8</figureCitation> ) and also in <taxonomicName id="4C5792FDFFB63D18DC3EF8F7FC0AF85C" authorityName="Aubrechtová & Korn" authorityYear="2022" box="[832,1001,1907,1933]" class="Cephalopoda" family="Lituitidae" genus="Lituites" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Orthocerida" pageId="67" pageNumber="68" phylum="Mollusca" rank="species" species="kruegeri"> <emphasis id="B923356CFFB63D18DC3EF8F7FCBBF85C" box="[832,856,1908,1933]" italics="true" pageId="67" pageNumber="68">L.</emphasis> cf. <emphasis id="B923356CFFB63D18DCF8F8F0FC0AF85C" box="[902,1001,1907,1933]" italics="true" pageId="67" pageNumber="68">kruegeri</emphasis> </taxonomicName> . Some specimens of <taxonomicName id="4C5792FDFFB63D18DB92F8F7FA9AF85F" authorityName="Wahlenberg" authorityYear="1818" box="[1260,1401,1908,1934]" class="Cephalopoda" family="Lituitidae" genus="Lituites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="67" pageNumber="68" phylum="Mollusca" rank="species" species="perfectus"> <emphasis id="B923356CFFB63D18DB92F8F7FA9AF85F" box="[1260,1401,1908,1934]" italics="true" pageId="67" pageNumber="68">L. perfectus</emphasis> </taxonomicName> have a large conch diameter ( <quantity id="4CAF449BFFB63D18DD6FF814FD80F860" box="[529,611,1943,1969]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="67" pageNumber="68" unit="mm" value="39.0">39 mm</quantity> in the <typeStatus id="54EC57DCFFB63D18DDCFF814FCEDF860" box="[689,782,1943,1969]" pageId="67" pageNumber="68" type="neotype">neotype</typeStatus> MB.C.30544) and a low WER ah ( <quantity id="4CAF449BFFB63D18DBEAF814FB34F860" box="[1172,1239,1943,1969]" metricMagnitude="-2" metricUnit="m" metricValue="4.826" pageId="67" pageNumber="68" unit="in" value="1.9">1.9 in</quantity> MB.C.30544 and MB.C.30545) as <taxonomicName id="4C5792FDFFB63D18DEBCF838FDC7F805" authorityName="Remele" authorityYear="1890" box="[450,548,1978,2004]" class="Cephalopoda" family="Lituitidae" genus="Lituites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="67" pageNumber="68" phylum="Mollusca" rank="species" species="fallax"> <emphasis id="B923356CFFB63D18DEBCF838FDC7F805" box="[450,548,1978,2004]" italics="true" pageId="67" pageNumber="68">L. fallax</emphasis> </taxonomicName> but the former species differs in the shell ornament of the coiled part; it consists of growth lines and rectiradiate annuli or ribs instead of sharp lirae. With only the <typeStatus id="54EC57DCFFB13D1FDA6EFB08FA94FB74" box="[1296,1399,1163,1189]" pageId="68" pageNumber="69" type="holotype">holotype</typeStatus> MB.C.11672.2 at hand, it is therefore not possible to confidently synonymise <taxonomicName id="4C5792FDFFB13D1FDB23FB2CFB22FB19" authorityName="Remele" authorityYear="1890" box="[1117,1217,1198,1224]" class="Cephalopoda" family="Lituitidae" genus="Lituites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="68" pageNumber="69" phylum="Mollusca" rank="species" species="fallax"> <emphasis id="B923356CFFB13D1FDB23FB2CFB22FB19" box="[1117,1217,1198,1224]" italics="true" pageId="68" pageNumber="69">L. fallax</emphasis> </taxonomicName> with any other species of the genus <taxonomicName id="4C5792FDFFB13D1FDECEFB51FDEBFB3A" authorityName="Lasaulxii Remele" authorityYear="1890" box="[432,520,1234,1259]" class="Cephalopoda" family="Lituitidae" genus="Lituites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="68" pageNumber="69" phylum="Mollusca" rank="genus"> <emphasis id="B923356CFFB13D1FDECEFB51FDEBFB3A" box="[432,520,1234,1259]" italics="true" pageId="68" pageNumber="69">Lituites</emphasis> </taxonomicName> . </paragraph> <caption id="DF28B9F6FFB13D1FDFC3FC35FCFFFBEA" ID-DOI="http://doi.org/10.5281/zenodo.6341370" ID-Zenodo-Dep="6341370" httpUri="https://zenodo.org/record/6341370/files/figure.png" pageId="68" pageNumber="69" startId="68.[189,232,950,976]" targetBox="[295,1295,272,906]" targetPageId="68"> <paragraph id="8BE8E97EFFB13D1FDFC3FC35FCFFFBEA" blockId="68.[189,1398,950,1083]" pageId="68" pageNumber="69"> <emphasis id="B923356CFFB13D1FDFC3FC35FEC0FC01" bold="true" box="[189,291,950,976]" pageId="68" pageNumber="69">Fig. 46.</emphasis> <taxonomicName id="4C5792FDFFB13D1FDE4CFC34FD74FC00" authority="Remele, 1890" authorityName="Remele" authorityYear="1890" box="[306,663,950,977]" class="Cephalopoda" family="Lituitidae" genus="Lituites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="68" pageNumber="69" phylum="Mollusca" rank="species" species="fallax"> <emphasis id="B923356CFFB13D1FDE4CFC34FE3FFC01" box="[306,476,950,976]" italics="true" pageId="68" pageNumber="69">Lituites fallax</emphasis> Remelé, 1890 </taxonomicName> from the Upper Grey Orthoceratite Limestone. Lectotype MB.C.11672.2 (Ramann, Kgl. Forstakademie Eberswalde Coll.) from Heegermühle (Brandenburg, Germany); photograph (with dorsal, lateral and ventral views) and reproduction of the illustration by Remelé (1890: pl. 5 fig. 1). Scale bar units = 1 mm. </paragraph> </caption> <paragraph id="8BE8E97EFFB13D1FDFC3FA91FE32F9D1" blockId="68.[189,1399,1298,1536]" pageId="68" pageNumber="69"> Remelé (1890: pl. 1 fig. 3) assigned a second specimen (MB.C.11673.2) from coeval strata to <taxonomicName id="4C5792FDFFB13D1FDA60FA90FEE3FA81" authorityName="Remele" authorityYear="1890" class="Cephalopoda" family="Lituitidae" genus="Lituites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="68" pageNumber="69" phylum="Mollusca" rank="species" species="fallax"> <emphasis id="B923356CFFB13D1FDA60FA90FEE3FA81" italics="true" pageId="68" pageNumber="69">Lituites fallax</emphasis> </taxonomicName> ? but in this specimen nothing of the coiled part is preserved and hence its correspondence to the <typeStatus id="54EC57DCFFB13D1FDF95FADAFEB1FAA2" box="[235,338,1369,1395]" pageId="68" pageNumber="69" type="holotype">holotype</typeStatus> cannot be validated. <bibRefCitation id="EFC6948FFFB13D1FDD28FADAFD12FAA2" author="Holm G." box="[598,753,1369,1395]" pageId="68" pageNumber="69" pagination="736 - 780" refId="ref47009" refString="Holm G. 1891. Om mynningen hos Lituites Breyn. Geologiska Foreningen i Stockholm Forhandlingar 13: 736 - 780. https: // doi. org / 10.1080 / 11035899109445850" type="journal article" year="1891">Holm (1891)</bibRefCitation> and <bibRefCitation id="EFC6948FFFB13D1FDC51FADAFC2CFAA5" author="Sweet W. C." box="[815,975,1369,1396]" pageId="68" pageNumber="69" pagination="1 - 176" refId="ref49478" refString="Sweet W. C. 1958. The Middle Ordovician of the Oslo region, Norway. 10. Nautiloid cephalopods. Norsk Geologisk Tidsskrift 38: 1 - 176." type="journal article" year="1958">Sweet (1958)</bibRefCitation> argued that the specimen probably represents <taxonomicName id="4C5792FDFFB13D1FDE40FAFFFDABFA47" baseAuthorityName="Holm" baseAuthorityYear="1891" box="[318,584,1404,1430]" class="Cephalopoda" family="Lituitidae" genus="Trilacinoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="68" pageNumber="69" phylum="Mollusca" rank="species" species="discors"> <emphasis id="B923356CFFB13D1FDE40FAFFFDABFA47" box="[318,584,1404,1430]" italics="true" pageId="68" pageNumber="69">Trilacinoceras discors</emphasis> </taxonomicName> . Indeed, this specimen is similar in ornament, expansion angle and general adult size to <taxonomicName id="4C5792FDFFB13D1FDED6FA23FDFAFA68" baseAuthorityName="Holm" baseAuthorityYear="1891" box="[424,537,1439,1465]" class="Cephalopoda" family="Lituitidae" genus="Trilacinoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="68" pageNumber="69" phylum="Mollusca" rank="species" species="discors"> <emphasis id="B923356CFFB13D1FDED6FA23FDFAFA68" box="[424,537,1439,1465]" italics="true" pageId="68" pageNumber="69">T. discors</emphasis> </taxonomicName> . However, the specimen is still chambered and its whorl profile is circular. Its identity is thus left unresolved herein; the <typeStatus id="54EC57DCFFB13D1FDC7CFA41FC8AFA0D" box="[770,873,1474,1500]" pageId="68" pageNumber="69" type="holotype">holotype</typeStatus> remains the only specimen unequivocally attributable to <taxonomicName id="4C5792FDFFB13D1FDE16FA65FE29F9D1" authorityName="Remele" authorityYear="1890" box="[360,458,1510,1536]" class="Cephalopoda" family="Lituitidae" genus="Lituites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="68" pageNumber="69" phylum="Mollusca" rank="species" species="fallax"> <emphasis id="B923356CFFB13D1FDE16FA65FE29F9D1" box="[360,458,1510,1536]" italics="true" pageId="68" pageNumber="69">L. fallax</emphasis> </taxonomicName> . </paragraph> </subSubSection> </treatment> </document>