Taxonomic and Zoogeographic Study of the Japanese Phygadeuontinae (Hymenoptera, Ichneumonidae), with Descriptions of 17 New Species Author Watanabe, Kyohei Kanagawa Prefectural Museum of Natural History, 499 Iryuda, Odawara, Kanagawa 250 – 0031, Japan watanabe-k@nh‛kanagawa-museum‛jp text Bulletin of the Kanagawa Prefectural Museum (Natural Science) 2021 2021-03-30 50 55 136 https://doi.org/10.32225/bkpmnh.2021.50_55 journal article 300009 10.32225/bkpmnh.2021.50_55 70224806-ca38-47ca-bd8a-63da3ac532f9 2189-6720 12774639 DD695C62-DC2F-4D32-8D33-52648E28661D Hemiteles maculipterus sp. nov. (New SJN: Hanemon-mame-togari-himebachi) ( Figs 42A–E , 63I , 64D , 66L ) Etymology . The specific name is from the fore wing with clouded maculation. Type series . Holotype : KPM-NK 55025 , F, JAPAN , Shizuoka Pref. , Kawanehonchou Town , Mt. Yamainudan , 14. VI. 2008 , K. Watanabe leg. Paratypes : JAPAN , KPM-NK 55026 , F, ditto ; KPM-NK 55027 , F, Kagoshima Pref. , Yakushima Is. , Arakawa , 4–7. VI. 2007 , T. Yamauchi leg. (MsT) ; KPM-NK 55028 , F, ditto, 22. VII. – 22. VIII. 2006 ; KPM-NK 55029–55031 , 3 F, ditto, 29. VII. – 25. VIII. 2007 . Description . Female (n=7). Body length 3.7–5.2 (HT: 5.2) mm. Body matt and densely coriaceous, covered with silver setae. Head. Clypeus 2.0 times as wide as maximum length, its anterior margin rounded. MSL 1.0–1.1 (HT: 1.0) times as long as BWM. OOL almost as long as POL. OOL and POL longer than OD. Occipital carina complete, its lower end joined with hypostomal carina distant from mandibular base. Upper tooth of mandible slightly longer than lower tooth. Base of mandible flat. Antenna with 26–30 (HT: 30) flagellomeres. FL III 3.3 times as long as maximum depth in lateral view. Mesosoma. Lateral part of pronotum covered with minute longitudinal striae ventrally. Epomia indistinct. Notaulus short, fading out in front of the center of mesoscutum. Scutellum without a lateral longitudinal carina except for base. Mesopleuron covered with minute longitudinal striae ventrally except for coriaceous speculum and posterior part. Posterior transverse carina of mesosternum largely absent in front of mid coxae. Metapleuron without a juxtacoxal carina. Anterior transverse carina and posterior transverse carina of propodeum complete ( Fig. 64D ) or lateral part of the latter sometimes partly indistinct. Lateral longitudinal carina absent except for distinct anterior section ( Fig. 64D ). Lateromedian longitudinal carina of propodeum complete ( Figs 42E , 64D ). Area basalis distinct ( Fig. 64D ). Area superomedia distinct, transversely elongate, its length almost as long as area basalis ( Fig. 64D ). Postero-lateral corner of area dentipara weakly projected. Area petiolaris strongly elongated ( Fig. 64D ). Median and posterior parts of propodeum irregularly rugulose except for transverse rugae on area petiolaris. Length of fore wing 4.0–4.4 (HT: 4.1) mm. Areolet absent ( Fig. 42D ). Vein 2m-cu of fore wing with a single bulla. Nervellus inclivous, intercepted behind the middle ( Fig. 42D ). Hind femur 4.7–4.9 (HT: 4.7) times as long as maximum depth in lateral view. Hind TS I: II: III: IV: V = 2.0: 0.9–0.95 (HT: 0.9): 0.65–0.7 (HT: 0.65): 0.3: 0.4. Tarsal claws simple. Metasoma. T I 2.4–2.7 (HT: 2.7) times as long as maximum width, longitudinally and obliquely striate, sometimes with a shallow median longitudinal concavity posteriorly. Median dorsal carina of T I absent. Dorsolateral carina of T I indistinct. T II to T VI wider than long, finely and densely punctate. Ovipositor sheath 0.75–0.8 (HT: 0.75) times as long as hind tibia. Ovipositor straight except for slightly upcurved apex, without a nodus and with weak ventral teeth ( Fig. 66L ). Coloration ( Figs 42A–E ). Body (excluding wings and legs) black to blackish brown. Scape, pedicel, and base of FL I reddish brown to brown. Mandible yellowish brown except for darkened apex and base. Palpi and tegula whitish yellow to yellowish brown. Apex of T I, T II, and base of T III reddish yellow. Posterior margins of T III to T VI narrowly tinged with reddish brown. Posterior part of T VII white. T III to T VII sometimes tinged with brown. Membranous part of metasomal sternites whitish yellow. Ovipositor yellowish brown. Wings hyaline, area behind of pterostigma largely clouded. Veins and pterostigma brown to dark brown. Fore and mid coxae and all tibial spurs, trochanters and trochantelli whitish yellow to yellow. Fore and mid femora, tibiae, and tarsi yellowish brown to dark brown. Hind coxa and base of femur reddish brown. Hind femur except base, tibia except for base, and tarsus blackish brown. Base of hind tibia tinged with whitish yellow. Male. Unknown. Distribution . Japan (Honshu and Yakushima Is.). Bionomics . Unknown. Remarks . This species resembles H. similis ( Gmelin, 1790 ) and H. rubropleuralis Kiss, 1929 , but it can be distinguished by the following combination of character states: the fore wing with clouded area (absent in H. similis ); FL III 3.3 times as long as maximum depth in lateral view (2.5–3.0 in H. rubropleuralis ); mesosoma without red area (usually with red area in H. similis ).