Taxonomic revision of the Australian Eucarpiini (Hemiptera: Fulgoromorpha: Cixiidae) with the description of nine new species
Author
Löcker, Birgit
Author
Fletcher, Murray J.
Author
Gurr, Geoff M.
text
Zootaxa
2010
2425
1
31
journal article
10.5281/zenodo.194600
e82c67cc-8d49-49bd-a0cd-d64b2bda9aec
1175-5326
194600
Dilacreon
(
Dilacreon
)
granulinervis
(
Muir, 1913
)
,
comb. nov.
(
Figs 4
,
14
)
Ptoleria granulinervis
Muir, 1913
: 256
.
Eucarpia granulinervis
(Muir)
,
Fennah 1980
: 238
.
Dilacreon (Dilacreon) granulinervis
(Muir)
,
comb. nov
.
Notes:
Eucarpia granulinervis
is hereby transferred into
Dilacreon
(
Dilacreon
) Fennah
, based on the characters of the wings, chaetotaxy of the hind legs and shape and colour of the head as given in
Fennah (1980)
. The transfer of this species to
Dilacreon
places this genus in the Indonesian fauna for the first time.
Types
.
Lectotype
3 (here designated, examined),
INDONESIA
: Larat, Timor Laut Islands (792) (F. Muir) (BPBM).
Paralectotype
,
INDONESIA
: 1 Ƥ, same data,
xii.1907
(BPBM).
Notes: The original description does not mention the designation of a
holotype
, therefore these specimens are regarded as
syntypes
. A
lectotype
is designated to provide a diagnostic reference for the species.
Other material examined.
AUSTRALIA
, Qld
: 33, 1Ƥ, St. Pauls, Moa Island,
10.11S
142.26E
, at light,
10.–18.ii.1986
(K. Houston & E. Hamacek) (
QDPI
); 13, ‘Eclectus’, Iron Range,
12º45’46”S
143º17’10”E
,
20 m
,
12.vii.1997
(G. & A. Daniels) (
UQIC
); 1 3,
Batavia
Downs,
12.41S
142.41E
, flight intercept trap,
22.vi.– 23.viii.1992
(P. Zborowski & J. Cardale) (
ANIC
); 3 3, same data, malaise trap (
ANIC
); 17 3, 22 Ƥ,
5 km
S
Batavia
Downs,
12.41S
142.41E
, malaise trap,
24.v.–17.vi.1993
(P. Zborowski & I. Naumann) (
ANIC
); 1 Ƥ, same data,
4.iv.–24.v.1993
(P. Zborowski & A. Rosch) (
ANIC
);
5 3, 12
Ƥ, same data,
18.vi.–22.vii.1992
(P. Zborowski & E.S. Nielson) (
ANIC
); 1 Ƥ, same data,
8.iii.–4.iv.1993
P. Zborowski) (
ANIC
); 1 3, same data,
12.40S
142.41E
, flight intercept trap,
18.vi.–22.vii.1992
(P. Zborowski & E.S. Nielson) (
ANIC
); 5 3, 7 Ƥ, Heathlands,
11.45S
142.35E
, flight intercept trap,
18.viii.–ix.1992
(P. Zborowski & L. Miller) (
ANIC
); 1 Ƥ, Lockerbie, Cape York, 6.-1-.
vi.1969
(G.B. Monteith) (
UQIC
); 1 Ƥ, Gordon Creek, Claudie River District,
12º42’S
143º17’E
, mv lamp,
6.xii.1986
(G. Daniels & M.A. Schneider) (
UQIC
); 1 Ƥ, Mount Lamond, Iron Range,
28.vi.1982
(G. Daniels & M.A. Schneider) (
UQIC
).
Colour.
Head, pro- and mesonotum light brown with concolorous carinae (some specimens with pale yellow carinae); face with three whitish spots along frontoclypeal suture as in
Fig. 4
E. Forewing hyaline light brown, sometimes with darker markings, veins concolorous with cells (some specimens with pale carinae); tubercles darker than veins and cells; pterostigma light brown. Fore- and midlegs light brown with mid brown transverse stripes as in
Fig. 4
A.
Hind
legs light brown, not striped, but with darkened areas around the apical teeth. Abdominal sternites light brown to mid brown.
Morphology.
Body length: 3
3.9–4.7 mm
; Ƥ 4.0–
4.8 mm
.
Head: Vertex 1.9–2.4 times wider than long; with distinct median carina. Frons 0.8–1.0 times as long as wide; frons invisible in dorsal view. Rostrum slightly surpassing hind coxae.
FIGURE 4.
Dilacreon (Dilacreon) granulinervis
: A habitus lateral; B forewing; C hindwing; D dorsum; E face; F head, anterodorsal view. Scalebar 1 mm applies to figure A.
Thorax: Mesonotum with lateral carinae sinuate as in
Fig. 4
D. Forewing 2.8–3.0 times longer than wide; with about 23–28 tubercles on costa; Sc+R fused, forming common stem Sc+R, M emerging separately from basal cell; Sc+R forking basad, at same level or distad of fork CuA1+CuA2; position of crossvein R-M at same level as fork MA-MP.
Male genitalia: Anal tube (
Figs 14
C, D), more or less symmetrical in dorsal view, apical lobes with pointed tip as in
Fig. 14
D. Genital styles as in
Figs 14
E, F, inner side of basal arm without tooth. Ventromedian process of pygofer triangular as in
Fig 14
E. Aedeagus as in
Figs 14
A, B. Phallotheca ventrally with one triangular and one ovoid shaped, flattened ridge; right laterally with short spine (b) inserted near apex of aedeagus; left laterally with longer spine (d) directed cephalad and very thin spine (f) directed caudad. Flagellum dorsally with short spine (a); right laterally with longer spine (c) and left laterally with short spine (e).
Remarks.
This species can be distinguished from the other two Australian species by the presence of a pointed tip on the apical lobes of the anal tube (rounded in
D.
(
D.
)
akethe
and
D.
(
D.
)
ispi
), by the presence of a triangular and an ovoid shaped, flattened ridge on the ventral side of the aedeagus (absent in
D.
(
D.
)
akethe
and
D.
(
D.
)
ispi
) and by the absence of a small tooth on the inner side of the basal arm of the genital style.
Dilacreon
(
D.
)
granulinervis
was previously known only from
Indonesia
but it seems to be a common species in North Queensland. Thirty-six males and
47 females
have been found in the Australian material representing the first record of the species in
Australia
.