Revision of the Ipsviciidae of the Late Triassic of Queensland (Hemiptera Cicadomorpha: Scytinopteroidea) Author Lambkin, Kevin J. text Zootaxa 2020 2020-10-14 4860 4 503 520 journal article 8250 10.11646/zootaxa.4860.4.2 7fc75baa-2e48-490b-9a6e-256378b4531e 1175-5326 4414254 C5095CD0-1DF5-4BE9-9B31-0CDB9443CF7F Ipsvicia Tillyard, 1919 Ipsvicia Tillyard, 1919a: 878 . Type species. Ipsvicia jonesi Tillyard, 1919 , by original designation. Emended diagnosis. Tegmen c . 3 times longer than wide; surface sculpture complex, comprising numerous variably shaped patches of fine tubercles each encircled by a smooth, very narrow margin, the patches set in a coarser tuberculate/punctate groundmass, a little finer apically; costal margin distinctly convex; basal cell short and broad, not extending to midpoint of PCu; R evenly convex, parallel with costal margin, the basal R cell therefore of about equal width for entire length; two r-m ; CuA strongly angulate at m-cua ; base of 1A slightly upcurved. Description. Tegmen c . 14–22 mm long; marginal membrane only slightly widened postero-apically; costal fracture fairly upright; bSc variable convex; dSc very weakly developed, running along middle of costal space; R with 4–6 mostly simple post 1r-m branches; M+CuA stem a little variable in length, extending just beyond ½ length of PCu; apical section of CuA directed posteriorly, straight or slightly convex; clavus extending to c . 0.6 tegmen length; 1A about as long as PCu+1A. Hind wing: size variable in proportion to variable size of the individual, maximum length c . 13.8 mm , maximum width c . 5.9 mm in measurable specimen; R, M, CuA and CuP very closely approximated basally, CuA and CuP apparently without a common stem; R, M and CuA then more or less parallel, directed towards apical margin, gradually diverging apically, apex of CuA upcurved and forming an apical loop with R and M enclosing a small network of terminal short branches and crossveins; a long, backwardly oblique m-cua (the single cua-cup illustrated by Tillyard (1923) and Evans (1963) not detected); CuP simple, very long, sinuous, its apex broadly separated from that of CuA and PCu; both PCu and 1A with thickened bases, both straight basally and sinuous apically, CuP and PCu approximating each other at their mid-points; anal area noticeably crinkled; marginal membrane well developed. Notes. The hind wing venation is noteworthy for the close alignment and apical looping of R, M and CuA, and the long, simple and sinuous CuP, PCu and 1A. The other known hind wings of the Scytinopteroidea are those of several species of Scytinoptera Handlirsch, 1904 , illustrated by Becker-Migdisova (1948 , figs. 2, 33, 39–43). They apparently lack a marginal membrane, and their venation is much simpler than that of Ipsvicia , with M and CuA directed postero-apically rather than apically, R, M and CuA without apical looping, CuA and CuP with a common stem, CuP, PCu and 1A almost straight rather than sinuous, and without the long, backwardly oblique m-cua . Indeed, as noted by Evans (1956) , the venation of Ipsvicia was unlike any of the Palaeozoic or Mesozoic homopteran hind wings known at that time (see also those illustrated by Becker-Migdisova 1961 ), and it differs from that of the extant families of Cicadomorpha (see figures in Kramer 1950 ). It might be observed, however, that the form of R, M and CuA bears some resemblance to that of Heteroptera: Nepomorpha ( Popov 1971 , plate 52).