A revision of Canarium L. (Burseraceae) in Madagascar Author Daly, Douglas C. Author Raharimampionona, Jeannie Author Federman, Sarah text Adansonia 2015 2015-12-31 37 2 277 345 https://www.mendeley.com/catalogue/2522e05a-0ab0-3e53-96aa-d45147a2ca6c/ journal article 10.5252/a2015n2a2 1639-4798 5208903 Canarium indistinctum Daly, Raharim. & Federman , sp. nov. ( Figs 6 ; 16 ). Medium-sized trees, leaves 1-3-jugate, stipules 2-3 mm long, the stipular scar 1-1.5 mm long, lateral leaflet base usually symmetric, petiole and rachis with only sparse to scattered hairs; distinguished from C. elegans Daly, Raharim. & Federman , sp. nov. by the stipules 8-24 (vs 4-8 in C. elegans , sp. nov. ) mm from petiole insertion, the lateral leaflets elliptic or less often oblong-elliptic or oblanceolate (vs ovate to slightly oblong-ovate), the fruit pedicel 4-10 (vs 3-4) mm long, and the fruit 3.2-5.2 × 1.7-2.6 cm (vs 2.8-3.5 × 1.7-1.9 cm ) and (ob)ovoid to oblong-ellipsoid (vs (oblong-)obovoid and trigonous). TYPUS . — Madagascar . Toamasina , path from “Pépinière” of Ambatondradama , Masoala Peninsula , Mahalevona , Maroantsetra , 375- 1011 m elev., 15°17’22”S , 50°00’58”E , 8.X.1997 , C. Birkinshaw , H. Ralimanana & R. Ranaivojaona 494 (holo-, NY !; iso-, MO !, TAN ). DISTRIBUTION AND ECOLOGY. — Known only from the type collection. So far as it is known, this species is a relatively small rainforest tree found at c. 950 m elevation near Maroantsetra on the Masoala Peninsula. Fruiting Jul-Oct . ETYMOLOGY. — The specific epithet refers to the inconspicuous venation that characterizes this species. DESCRIPTION Trees to at least 15 m tall. Resin white when dry. Leaves 11- 21 cm long, 1-3-jugate; petiole 1.8-4.7 cm long, petiole and rachis with a few scattered flexuous to ascending white hairs to 0.3 mm long; stipules 4-8 mm from petiole insertion, c. 2 mm long, obovate and constricted at base, folded laterally, with dense appressed white hairs to 0.15 mm long, the scar 1-1.5 mm long and raised; basal petiolules 5-8 mm long, other laterals 4-12 mm long, the terminal one 12-30 mm long, petiolules deeply canaliculate, pulvinuli not conspicuous; basal leaflets 32-85 × 23-42 cm , broadly ovate to very broadly elliptic, other laterals 60-12.2 × 3.0- 4.4 cm , ovate to slightly oblong-ovate, terminal leaflet 6-10.4 × 3.2-4.6 cm , (slightly) (ob)ovate; leaflet apex abruptly and very broadly short-acuminate, the acumen 1-5 mm long; lateral leaflet base usually symmetric, sometimes slightly oblique, obtuse and slightly attenuate; margin flat; leaflets drying brown, coriaceous, dull; secondary vein framework festooned-brochidodromous, secondaries in 7-13 pairs, spreading or almost straight, spacing decreasing toward apex and base, the angle slightly increasing toward base, perpendicular epimedial tertiaries present (these reticulating distally), intercostal tertiaries random-reticulate and admedially branched, quaternaries random-reticulate; on abaxial side the midvein broadly prominent, rest of veins narrowly prominulous, with scattered appressed, thick, white hairs to 0.5 mm long on midvein and secondaries, plus a few on rest of surface; on adaxial surface the midvein narrowly prominulous but sunk in a groove, secondaries narrowly prominulous or flat, rest usually flat, sometimes broadly prominulous, surface glabrous except a few scattered ascending white hairs at base. Infructescence 5.5-11 cm long × 2.5 mm diam above base, axes with dense, flexuous, fine pale hairs to 0.15 mm long and sparse capitate glands; fruiting pedicel 3.7-4.2 × 3.7-4.2 mm , clavate, fruiting calyx 4-6 mm long, continuous with the pedicel, calyx lobes distinct, 2-3 mm long, rounded-triangular, nearly patent. Fruit 2.8-3.5 × 1.7- 1.9 cm [ c. 4 × 2.5 cm when fresh], (oblong-)obovoid and trigonous, the apex rounded to slightly acuminate, the base acute to rounded; surface with dense, small, slightly raised, ferrugineous lenticels, also provided with descending, flexuous, white hairs to 0.4 mm long among lenticels, glabrescent. FIG. 18. — Canarium lamianum Daly, Raharim. & Federman , sp. nov. : A , flowering branchlet; B , staminate flower with bracteole; C , adaxial view of petal; D , sta- mens and ovariodisk; E , portion of infructescence; F , fruiting branchlet; G , leaf with detail of stipules ( G’ ) and leaflet margin ( G’’ ); A -D , Daly et al. 13065 (NY); E , Treuer et al. 44 (NY); F , McPherson & van der Werff 16378 (MO); G , Antilahimena et al. 663 (NY). Scale bars: A, E-G, 2 cm; B, 3 mm; C, D, 2 mm; G’, G’’, 5 mm. NOTES Canarium indistinctum , sp.nov. is distinguished from C. elegans , sp. nov. in the discussion under the latter species.