New species of Dugdaleiella, gen. nov., Kozloviella, gen. nov., and Pfitzneriella Viette from upper elevation Andes of Ecuador and Peru (Lepidoptera: Hepialidae) Author Grehan, John R. Author Mielke, Os. G. C. text Zootaxa 2018 2018-10-08 4497 1 1 28 journal article 29235 10.11646/zootaxa.4497.1.1 d5b94fdb-db02-4530-85aa-12d9757e3313 1175-5326 1451056 890E26F1-1B03-4F13-9299-A5E8BED8439D Pfitzneriella Viette, 1951a ( Figs 2–7 , 9b–9f , 10b–10h , 11b–11f , 12, 13b–13e, 14a–14e , 15b, 15c, 16b– 16g , 18–24b ) Type species: Triodia remota Pfitzner, 1906 by original designation. Diagnosis . The presence of a rounded and setose distal posterior corner of the valva and a lateral pointed or blunt saccus spur distinguish Pfitzneriella from all other Hepialidae that we have been able to examine. Pseudotegumen is not fused mesally, in contrast to Dugdaleiella gen. n. and Kozloviella gen. n . The ventral unfused condition of the pseudotegumen is also present in many other Hepialidae . In the cibyrine clade and the southern Andean genera the valvae have a range of shapes that are either elongate or very short, but lacking the distinct outer angle of Pfitzneriella ; those of Dugdaleiella gen. n. are of similar width throughout; the valvae of Kozloviell a gen. n. are bifurcated. The paired posterior projections of the saccus contrast with the short rounded knobs of Druceiella (Grehan & Rawlins in press) and Gymelloxes Viette (Grehan & Mielke 2017) . Narrowly triangular projections also occur near the midline in most species of Dalaca Walker, 1856 ( Nielsen & Robinson 1983 ) but it is not certain as to whether they are on the posterior saccus or project from sternum VIII. Remarks . Viette (1951a) originally characterized Pfitzneriella as having slightly fusiform antennae, small head, rather long and thin labial palpi extending beyond the head, and epiphysis present. In addition, FW CuP (as Cu2) not reaching the middle of the wing, rounded apex of pseudotegumen, broad and upright valvae, saccus with a spur on each side, and fultura inferior (juxta) forming a large plate. Later, another taxon was established, Pfitzneriella similis Zukowski, 1954 which was never illustrated and apparently destroyed by the bombing of Hamburg during World War II (Zukowski 1954). Of the features listed by Viette (1951a) only the presence of a lateral spur on the saccus is close to being diagnostic for the genus, only if the blunt or rounded spur of P. titarenkoi sp . n . is included.