New fossil gastropod species (Mollusca: Gastropoda) from the upper Miocene of the Canary Islands (Spain)
Author
Martín-González, Esther
Author
Vera-Peláez, José Luis
Author
Castillo, Carolina
Author
Lozano-Francisco, M. Carmen
text
Zootaxa
2018
4422
2
191
218
journal article
30040
10.11646/zootaxa.4422.2.3
eec03b6a-265e-42a3-a24c-2768daeb791a
1175-5326
1251497
4182A649-56E2-448D-B130-06FE28DEC013
Patella tintina
Martín-González & Vera-Peláez
sp. nov.
Figure 5 A, B, C
1976
Patella ambroggii
Lecointre. Meco
: p. 88.
2001
Patella
cf.
caerulea
Linnaeus. Martín-González
et al.
: p. 52.
2001
Patella caerulea
Linnaeus. Castillo
et al.
: p. 68.
2008
Patella ambroggii
Lecointre. Meco
et al.
: p. 80, fig. 2.11.
2012
Patella ambroggii
Lecointre. Betancort Lozano
: p. 97, plate 6, figs. 4, 4a.
2015
Patella ambroggii
Lecointre. Meco
et al.
: p. 65, appendix 2, figs. a, b, c, d, g.
Type material.
Holotype: adult specimen, with broken apex, large size (L:
100.6 mm
; W:
85.2 mm
and H:
50.1 mm
), record number TFMCFO-3438/1. Paratypes: four well-preserved specimens, TFMCFO- 3535 (L:
78.6 mm
; A:
76.5 mm
; H:
25.7 mm
); TFMCFO-4601 (L:
52.7 mm
; W:
40.9 mm
; H:
12.6 mm
); TFMCFO-4 608 (L:
99 mm
; W:
97.2 mm
), and TFMCFO-6107 (L:
42.9 mm
; W:
33.8 mm
; H:
12.1 mm
).
Other material examined.
Three more specimens from Lanzarote, with numbers TFMCFO-3836, Punta Gorda and TFMCFO-6483, Punta Garajao; and 25 more from Fuerteventura (TFMCFO-3438, North of Agua Liques; TFMCFO-3718, Punta del Viento; TFMCFO-4593, Fuente Blanca; TFMCFO-4594, Barranco Gavioto; TFMCFO-4606, Playa del Valle; TFMCFO-4603, TFMCFO-5991, TFMCFO-6404, Bajas Amarillas; TFMCFO- 5992, Caletones Mansos; TFMCFO-6107, Aljibe de la Cueva; and TFMCFO-6578, Barranco de la Cruz). Biometric data for this material are shown in
Table 1
.
Type
locality.
North of Agua Liques
(
Fuerteventura
,
Canary Islands
); UTM: 28 R
573615 m
E
3118639 m
N
,
10 m
asl
. Sandstone deposit, approximately
50 cm
in thickness. Tortonian dating defined by the gastropod assemblage described in the geological and geographic context section above. The
paratypes
are from the localities:
Caletones Mansos
(TFMCFO-3535),
Barranco León
(TFMCFO-4608),
Punta
de Leonardo
(TFMCFO- 4601) and
Aljibe de la Cueva
(TFMCFO-6107), all in
Fuerteventura.
FIGURE 4.
Biochronostratigraphy distribution of upper Miocene species indicators.
TABLE 1.
Measurements of
Patella tintina
sp. nov.
| Record number |
La |
Lb |
L |
W |
H |
| TFMCFO-3438/1 |
100,6 |
85,2 |
| TFMCFO-3438/2 |
32,7 |
45,2 |
77,9 |
62,4 |
21,8 |
| TFMCFO-3535/1 |
36,4 |
42,3 |
78,6 |
76,5 |
25,8 |
| TFMCFO-3535/2 |
112,2 |
| TFMCFO-3535/3 |
41,7 |
| TFMCFO-3718 |
87,9 |
84,1 |
| TFMCFO-3836/1 |
18,1 |
27,7 |
45,8 |
12,7 |
| TFMCFO-3836/2 |
33,1 |
14,1 |
| TFMCFO-4593 |
17,1 |
22,2 |
39,4 |
28,8 |
10,1 |
| TFMCFO-4594/1 |
33,2 |
38 |
71,2 |
29,4 |
| TFMCFO-4594/2 |
32 |
41,5 |
73,5 |
53,1 |
24,3 |
| TFMCFO-4594/3 |
28,6 |
33,2 |
61,8 |
54,3 |
| TFMCFO-4594/4 |
63,0 |
57,1 |
| TFMCFO-4601 |
22,8 |
29,9 |
52,7 |
40,9 |
12,6 |
| TFMCFO-4603 |
21,6 |
16,6 |
| TFMCFO-4606 |
80,2 |
68,6 |
34,8 |
| TFMCFO-4608/1 |
36,7 |
43,3 |
79,9 |
75,1 |
35,7 |
| TFMCFO-4608/2 |
98,8 |
93,5 |
| TFMCFO-4608/3 |
99,1 |
97,2 |
| TFMCFO-5991 |
12,2 |
18,6 |
30,6 |
12,8 |
8,2 |
| TFMCFO-5992/1 |
37,7 |
30,6 |
| TFMCFO-5992/2 |
27,7 |
34,8 |
62,5 |
43,7 |
22,8 |
| TFMCFO-6107/1 |
6,9 |
9,5 |
16,4 |
13,2 |
3,2 |
| TFMCFO-6107/2 |
19,4 |
23,5 |
42,9 |
33,7 |
12,1 |
| TFMCFO-6404 |
23,4 |
31 |
54,4 |
34 |
13,3 |
| TFMCFO-6483 |
51,1 |
53,6 |
104,7 |
91,7 |
51,4 |
| TFMCFO-6578/1 |
28,6 |
36,6 |
65,2 |
48,7 |
25,3 |
| TFMCFO-6578/2 |
34,3 |
42,1 |
76,4 |
52,1 |
31,5 |
| Variables |
N |
Minimum |
Maximum |
Mean |
Standard error |
Standard desviation |
| La |
17 |
6,9 |
51,1 |
27,9 |
2,9 |
10,3 |
| Lb |
18 |
9,5 |
53,6 |
34,6 |
2,9 |
10,6 |
| L |
26 |
16,4 |
112,2 |
68,7 |
5,5 |
25,4 |
| W |
24 |
12,8 |
97,2 |
56,3 |
5,5 |
25,3 |
| H |
18 |
3,2 |
51,4 |
21,8 |
3,2 |
11,1 |
Etymology.
The name
tintina
refers to the metallic sound produced by this limpet shell when struck.
Description.
Large-sized shell (Lmax:
112 mm
; Wmax:
97.2 mm
; Hmax:
51.4 mm
), fragile, with thinner section in the apical region and thicker towards the base. Patelliform shape, with conical cap shape in adult phases, more flattened in juveniles. Shell base oval, wider in the posterior region than the anterior. Apex situated further towards the anterior region at nearly 2/3 from the posterior edge. Sculpture consists of six wide main ribs interspersed with a variable number of secondary and tertiary ribs in the anterior region; these become wider during ontogeny. In the posterior region, there are four strongly protruding main ribs, and a variable number of secondary radial ribs placed between the former. Main ribs have a rounded blunt profile. Intercostal spaces between posterior ribs are twice as wide as those in the anterior region, where the two central ribs lie very close to each other. In juvenile forms, the shell surface is covered with scales arranged radially, which are more clearly defined on the secondary ribs and disappear during ontogeny. Growth lines are clearly visible towards the lower middle part of the shell, well-defined and slightly sinuous, in some cases forming darker, circular bands seen also on the ventral side. Shell edge is sinuous and star-shaped. Mantle impression is not visibly apparent in any of the specimens, but marked in internal molds, being large, oval and open at one end, and located close to the base. Color is not preserved.
Remarks.
The new taxon has features resembling those of
Patella ambroggii
Lecointre, 1952
from the Moroccan Pliocene as illustrated by
Lecointre (1952)
. Therefore, several workers identified material from Neogene deposits of the
Canary Islands
as
Patella ambroggii
. After consulting photograps of the
holotype
of
P. ambroggii
(
Fig. 6
) deposited at the Muséum national d´Histoire naturelle in Paris, with record number R10111,
Patella tintina
sp. nov.
can be distinguished from
P. ambroggii
by having more and more densely arranged primary ribs. The base of
P. ambroggii
is more elongated and rippled, not fitting the pattern of four large, separate ribs in posterior position observed in the Canary species. In contrast to
Patella tintina
sp. nov.
,
P. ambroggii
has a secondary rib between the ribs. Moreover, growth lines on the shell surface are more pronounces in
P. ambroggii
and form concentric crests.
Patella tintina
sp. nov.
is higher and more conical, with blunter ribs, which give the shell a rounded appearance. The apex is more posteriorly located in the
Patella tintina
than in
P. ambroggii
.
Patella tintina
sp. nov.
differs in size and shell morphology from other smaller species from the Miocene of the Rhone valley such as
P. delphinensis
Fontannes, 1876
and
P. vindascina
Fontannes, 1876
(
Fontannes 1882, pl. 1, figs 9–10
). When compared to a juvenile form of
P. crassicosta
Rovasenda, 1897
(
in
Sacco 1897) from the middle Miocene of the Piedmont-Liguria basin (refigured in plate 3, fig. 5 of
Forli
et al.
2004
), the juvenile
Patella tintina
sp. nov.
can be distinguished by the posterior layout of its main ribs and the more triangular shell shape.
Patella alessia
Forli, 2004
, a Mediterranean Pliocene species, can be separated from
P. tintina
sp. nov.
by its greater number of main ribs, its lower height and its more star-shaped basal profile.
The large size of
P. tintina
sp. nov.
and its number and arrangement of ribs distinguish it from the three species of the
Patellidae
currently living in the
Canary Islands
:
Patella crenata
d'Orbigny, 1840
,
P. ulyssiponensis
Gmelin, 1791
and
P. piperata
Gould, 1846
. The large South African limpets
Cymbula oculus
(
Born, 1778
)
and
Cymbula granatina
(
Linnaeus, 1758
)
differ from
Patella tintina
sp. nov.
by having a finer and denser ribbing and a different rib layout on their anterior margin. The South African species also differ in their central apex location.
Patella tintina
sp. nov.
resembles the South African
Scutellastra longicosta
(
Lamarck, 1819
)
in its star-shaped margin and in having wide ribs, but the latter can be distinguished by its completely star-shaped aspect and different layout of the radial ribs, with four very wide and long ribs towards the posterior region, and three towards the anterior end.
Patella caerulea
Linnaeus, 1758
, is much smaller and very flat, differing notably in its ornamentation pattern.
Distribution.
Upper Miocene, Tortonian: Fuerteventura and Lanzarote (
Betancort Lozano 2012
); Gran Canaria (
Betancort Lozano 2012
;
Meco
et al.
2015
).