Omophorus (Sinomophorus) wallaCei: a new weevil from Borneo highlights the enigmatic Ethiopian-Oriental disjunct distribution (Coleoptera, Curculionidae, Molytinae) Author Tseng, Wei-Zhe Author Hsiao, Yun Author Hsu, Chen-Fu text Zootaxa 2018 2018-06-22 4438 3 588 596 journal article 29815 10.11646/zootaxa.4438.3.10 50118f26-9f46-40da-9739-2ca57527370c 1175-5326 1298162 439C6BBB-393A-470C-9332-383351BFAD2A Omophorus ( Sinomophorus ) wallacei sp. nov. ( Figs. 1–17 ) Type material. Holotype : ♂ , Mt. Trusmadi , Borneo , Malaysia , II. 2016 , C.- F. Hsu leg. ( ZRC ) . Paratypes : The same data as the holotype ( 1 ♂ in NMNS ; 1 ♂ in NTU; 1 ♂ in TARI ; 2 ♂ in ZRC , including one liquid preserved paratype ) . Description. Male ( holotype ). Measurements (in mm): SL: 9.6; RL: 2.9; RW: 1.1; PL: 2.5; PW: 4.4; EL: 7.7; EW: 6.9; AL: 2.0. Coloration: mainly dark brown dorsally, sparsely covered with yellowish brown scales; antennae reddish brown; legs dark brown; ventral side dark reddish brown, covered with dense, bifid, yellowish brown scales laterally and ventrally. Antennae and legs covered with hair-like yellow scales. Head: subspherical in dorsal view, forehead 0.9 × as wide as RW, somewhat convex laterally; surface densely punctate. Eyes large, broadly ellipsoidal, with length 1.4 × width. Rostrum straight, subparallel sided, somewhat widened basally and apically, with RL 1.1 × PL; surface densely punctate, with punctures mainly oblong, mesorostrum with medial sulcus, extending to occiput. Antennae: Scape inserted at basal half of rostrum, with length 14.1 × width, 1.0 × as long as funicle; funicle 7-segmented, I obconical, with length 1.8 × width, II obconical, somewhat shorter than I, with length 1.3 × width, III–VI subspherical, with length 0.5 × width respectively, 1.3 × as long as I, VII attached to club, with length 0.4 × width; club 3-segmented, clavate, extremely large, with length 1.8 × width, 0.7 × as long as funicle, suture between I and II indistinct while suture between II and III deep and obvious ( Figs. 3–4 ). Pronotum: subconical in dorsal view, strongly narrowed anteriorly, middle part strangled, with PL 0.6 × PW, PW 1.7 × width of anterior margin, side bisinuate, with paired transverse depression medially, surface lustrous metallic, punctate, with numerous spot-like punctures ( Fig. 1 ). FIGURES 1–4. Omophorus ( Sinomophorus ) wallacei sp. nov. : 1. Habitus, dorsal view. 2. Habitus, lateral view. 3. Head, lateral view. 4. Head, dorsal view. Scale bars: 1–2. 5.0 mm; 3–4. 1.0 mm. Scutellum: subtrapezoidal, rounded apically. Elytra: moderately convex laterally, ventral side slightly broadened, with obtuse humeral angle ( Fig. 5 ), tubercle developed basally in interstria III, and between interstria VII–IX, with EL 1.1 × width, EL 3.0 × PL, with paired indistinct tubercle between interstria IV–VI apically; surface lustrous metallic, densely punctate, punctures in various forms, widely scattered ( Figs. 1–2 ). Abdomen: ventrite II with length 0.1 × width, 1.2 × as long as III; III 1.1 × as long as IV, 1.1 × as long as V, width slightly narrowed apically ( Fig. 9 ), ventrite V concave apically. Tergite I–VI weakly sclerotized, tergite VII sclerotized, lateral margin strongly sclerotized, surface covered with dense hair-like scales ( Fig. 10 ), tergite VIII (pygidium) sclerotized, strongly sclerotized and emarginate apically, covered with dense hair-like scales ( Fig. 11 ). Legs: procoxae contiguous, mesocoxae 1.3 × as long as the distance between meso- and metacoxae; femora strongly broadened medially, with maximum width 1.7 × basal width, surface densely covered with hair-like scales; tibiae moderately straight, somewhat widened basally, with metatibial uncus developed; tarsi 5-segmented, tarsomere III rounded apically, tarsomere V divaricate, without paired ventral tooth. FIGURES 5–8. 5. Humeral angle of Omophorus ( Sinomophorus ) wallacei sp. nov. . 6. Humeral angle of Omophorus ( Pangomophorus ) biroi Voss, 1960 , holotype. 7. Vestiture of abdomen of Omophorus ( Sinomophorus ) wallacei sp. nov. 8. Vestiture of abdomen of Omophorus ( Pangomophorus ) biroi Voss, 1960 , holotype. Male genitalia and terminalia: sternite VIII semilunar, sclerotized, basal margin strongly sclerotized ( Fig. 13 ); spiculum gastrale widely emarginate apically, base little sclerotized ( Fig. 12 ). Aedeagus with AL 3.1 × width in dorsal view, somewhat curved at both sides laterally, lateral margin of pedon slightly recurved apically; endophallus with dense small teeth and very minute sclerotization ( Figs. 16–17 ); tegmen with ring, manubrium as long as parameroid lobes, parameroid lobes sclerotized medially, with sides somewhat sclerotized. ( Figs. 14–15 ). Female. Unknown. Diagnosis. This new species is similar to Omophorus ( Sinomophorus ) rongshu Wang, Alonso-Zarazaga, Ren & Zhang, 2011 , but can be easily distinguished by larger body size; distant eyes; shorter antennal club; rostrum somewhat broadened basally and apically; strongly bisinuate pronotal base; tubercle between interstria VII–IX present; aedeagus: larger in size (> 1.5 mm ), with broader, rounded apex, slightly recurved apically; tegmen with manubrium distinctly longer than parameroid lobes. FIGURES 9–13. Omophorus ( Sinomophorus ) wallacei sp. nov. : 9. Ventrites, ventral view. 10. Tergite VII, dorsal view. 11. Tergite VIII, dorsal view. 12. Spiculum gastrale, dorsal view. 13. Sternite VIII, dorsal view. Scale bars: 9. 5.0 mm; 10–13. 1.0 mm. FIGURES 14–17. Omophorus ( Sinomophorus ) wallacei sp. nov. : 14. Tegmen ring, dorsal view. 15. Lateral view. 16. Penis, dorsal view. 17. Lateral view. Scale bar: 1.0 mm. Etymology. The specific name is in honor of the renowned British naturalist, Alfred Russel Wallace, who is famous for his remarkable contributions on the theory of evolution and biogeography. Distribution. Malaysian Borneo.