Review of the Pacific leafhopper genus Hecaloidella (Hemiptera: Cicadellidae Deltocephalinae: Selenocephalini) with description of sixteen new species Author Xu, Deliang 0000-0003-0106-5570 Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, College of Plant Protection, Northwest A & F University, Yangling, Shaanxi 712100, China & 18710305867 @ 163. com; https: // orcid. org / 0000 - 0003 - 0106 - 5570 Author Webb, Michael D. 0000-0002-1312-6142 Department of Life Sciences, Natural History Museum, Cromwell Road, London, ME 15 6 QX, UK & mdwebb 04 @ gmail. com; https: // orcid. org / 0000 - 0002 - 1312 - 6142 mdwebb04@gmail.com Author Shang, Suqin 0000-0001-9376-0735 Biological Engineering Laboratory of Crop Diseases and Pests of Gansu Province, College of Plant Protection, Gansu Agricultural University, Lanzhou, Gansu 730070, China & shangsq @ gsau. edu. cn; https: // orcid. org / 0000 - 0001 - 9376 - 0735 shangsq@gsau.edu.cn Author Zhang, Yalin 0000-0002-1204-9181 Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, College of Plant Protection, Northwest A & F University, Yangling, Shaanxi 712100, China & yalinzh @ nwsuaf. edu. cn; https: // orcid. org / 0000 - 0002 - 1204 - 9181 yalinzh@nwsuaf.edu.cn text Zootaxa 2021 2021-03-22 4948 3 381 403 journal article 7470 10.11646/zootaxa.4948.3.4 6bfd846c-0304-4330-8941-95c406630230 1175-5326 4629100 8E0BF97A-D9E1-4F2E-BB63-82A6ED47E5DC H . nitida Osborn Figs. 1C , 3A–M H . nitida Osborn, 1934: 174 , fig 8; Zhang & Webb, 1966: 8, figs 2, 6, 10, 24, 99–104, 474; Shang & Zhang, 2012: 2–3 , fig 1. Redescription. Body length (including tegmina), male: 4.5–4.8mm ; female: 5.2–6.2mm . Head and thorax dorsally pale sordid yellow. Thorax and abdomen ventrally brown. Forewings brownish hyaline. Male genitalia with subgenital plate elongate triangular basally with short digitate apex, with a few short fine setae laterobasally. Connective with arms separated distally or joined loop-shaped, stem extended caudally into a long dorsally curved process, distally with a dorsal medial keel. Aedeagus very small, disassociated from connective; shaft very short and stout, terminating in a pair of short recurved hook-like processes, with a short to long spine-like process basally; gonopore apical on ventral surface. Material examined. Holotype : ♀ , Samoa , Upolu , Malololelei , 2000ft , 23.xi.1924 , P.A. Buxton and G.H. Hop- kins, NHMUK 010591769 ( BMNH ) . Paratypes : 1♂ , same data as holotype ( BMNH ) ; 10♂ 12♀ , Indonesia (Ma- luku), Utara and Tengah, C.J . Lomer, iv.1990 , on clove ( BMNH , NMW ) ; 3♂ , Papua New Guinea ( Sudest Island ), Mt Riu , viii-ix.1956, L.J. Brass , 5 th Archbold Expedition to New Guinea ( AMNH ) . Remarks. This species together with the three following species ( H . caudospina , H . nudapenis and H . pacifica ) is distinguished by its very small aedeagus disassociated from the connective. It is most similar to H . pacifica in having the connective strongly curved dorsally but differs in its paler colour and having the aedeagus with the basal spine narrower and the distal processes apical rather than subapical. The species occurs in two forms in the male genitalia, one from Samoa (single specimen, paratype ) with a short basal spine on the aedeagus ( Fig. 3I ) and another from other islands (see Material examined) with a longer spine ( Fig. 3J ).