Taxonomic notes on the genera Scatella and Scatophila (Diptera: Ephydridae) with a remark on Trixoscelis chilensis (Trixoscelididae) Author Zatwarnicki, Tadeusz Author Irwin, Anthony G. text Zootaxa 2018 2018-01-31 4377 1 91 109 journal article 30877 10.11646/zootaxa.4377.1.6 bf31986f-33c6-4a7b-8b58-736c352e54e4 1175-5326 1163298 CD3ED409-1877-4141-9AE3-ADAC1BE329BE Scatophila mesogramma (Loew) ( Figs. 18–23 ) Scatella mesogramma Loew, 1869 : 42 . Scatophila mesogramma : Sturtevant, 1923 : 12 (combination). Scatophila grisescens Frey, 1915 : 32 , new synonym . Scatophila infantina Becker, 1926 : 89 , synonymized by Zatwarnicki, 1987 : 282 . Type material examined: One female of the senior synonym, in good condition, was labelled: “ Scatophila mesogramma ♀ ”, "364", "R. I.", "Loew Coll.", "mesograma Lw.", " Type 11148", mounted directly, preserved in MCZ . The holotype female of the junior synonym, which is glued to a paper triangle, in moderate condition (left basal flagellomere missing, left wing broken at the base), labelled: empty gold round label; "Kasatschie 5. VII. 1909 " (handwritten), " Scatophila grisescens n. sp. ? Frey det." (name handwritten), " Holotypus " (printed, red), and is preserved in ZIN . FIGURES 18–19. Head with a part of thorax of Scatophila mesogramma Sturtevant, 1923 . 18. antero-oblique view. 19. lateral view (Sweden: Pitholmen; MZLU). Scale bar = 0.2 mm. Other material examined: CANADA : Northwest Territories : Chesterfield , 10. VII. 1950 , J. G. Chillcott , 4 ex ; Nova Scotia : East End , Sable Is. , 9. VI. 1967 , D. M. Wood , 2 ex, S. Harbour Bch . PG957946, 11. VIII. 1 983, J. R. Vockeroth, 26 ex; Quebec : Gt. Whale R., 23. VI.–24. VIII. 1949 , J. R. Vockeroth , 24 ex ( CNC ); Natashquwan , 23. VII. 1930 , W. J. Brown , 1 ex ( AMNH ) . USA : Oregon , Boiler Bay , 9–30. III. 1930 , J. Wilcox , 2 ♂ ; Massachusetts : Martha's Vineyard, Oak Bluffs, 13. VI. 1917 , C. W. Johnson , 1 ♀ ; New Jersey : Wildwood , 18. VII. 1908 , 1 ♀ (all ANSP ) . DENMARK : Frederikshavn , 30. VI. 1908 , Th. Mortensen , 2 ♀ ( ZMUC ) . GERMANY : Nordrhein-Westfallen : distr. Herford, Enger/ Bünde , 5. VII. 1994 , T. Zatwarnicki , 1 ♂ ( TZPC ) . POLAND : Wrocław-Leśnica , 10. V. 1985 , T. Zatwarnicki , 1 ♀ ( TZPC ) . RUSSIA : Karelia : Paanajärvi , R. Frey , 2 ♀ ( MZH ); Nenetsia : Pechora Bay ( 68°53'N , 53°34'E ), 25–26. VIII. 1994 , C. Hansson , 1 ♀ ( MZLU ) . SWEDEN : Norrbotten : Piteå, Pitholmen, 7. IX. 1965 , H. Andersson , 1 ♀ ( MZLU ) ; Skania , Borgeby , WSW Löddesnäs , 26. VII. 1966 , L. Hedström , 1 ♀ ( UUZM ). Type localities: Senior synonym was described from " Newport , Rhode Island " ( USA ) . Junior synonym was collected in Russia . " Jana-Land , Dorf Kasatschje am Fl. Jana " (= Kazachye , near Jana river ; coordinates 70°44' 2.3" N , 136° 13' 10.9" E ) . Diagnosis. The species is closely related to S. arenaria Cresson but is distinguished from other species by mostly pale, whitish wing, with a spotted pattern scarcely evident; gena broad, more than twice as wide as width of basal flagellomere ( Figs.18–19 ); mesonotum uniformly brownish-grey, frequently with small pale presutural spots ( Fig. 23 ). Male terminalia ( Fig. 20–22 ) with concave anterior margin of epandrium and triangular shape of anterior portion of the gonite, sclerotized part of the aedeagus with length twice the height. Distribution. Nearctic: Canada ( Northwest Territories , Nova Scotia , Quebec ), United States ( Maryland , Massachusetts , New Jersey , Oregon , Rhode Island ). Palaearctic: Belgium , Denmark , Finland , Germany , Great Britain , Poland , Russia (European Territory, Siberia), Sweden . Remarks. External features of the holotype of S. grisescens have been examined and clearly represent S. mesogramma , with which it is now formally synonymized.