The fast-running flies (Diptera, Hybotidae, Tachydromiinae) of Singapore and adjacent regions Author Grootaert, Patrick Author Shamshev, Igor V. text European Journal of Taxonomy 2012 2012-01-06 5 1 162 journal article 10.5852/ejt.2012.5 e809cd3a-36bb-413d-9f19-70837d7db34b 3715167 B69F002A-C1A0-439D-9477-62BFA87DEAD7 Stilpon singaporensis sp. nov. Figs 229-231 Diagnosis Species with brown thorax; antenna brownish; wing distinctly brownish infuscate, R 4+5 strongly curved towards costa in apical part; abdominal tergites 1 and 2 modified. Description Male LENGTH. Body 1.2 mm , wing length 1.1 mm . HEAD. Black. Occiput subshining, finely greyish pollinose. Two moderately long vertical bristles present. Frons linear with sides nearly parallel, rather broad, entirely tomentose, subshining. Ocellar tubercle with minute setulae. Antenna brownish (in paler specimens postpedicel somewhat paler). Palpus yellow, with strong, black subapical bristle. THORAX. Brown. Scutum entirely tomentose. Postpronotal bristle long, inclinate. Dorsocentral bristles in multiple rows, complete posteriorly. Acrostichal bristles 2-serial, complete posteriorly. LEGS. With fore tibia and tarsus (tarsomere 5 darker), mid femur in apical part, hind femur (except narrow basal space), mid and hind tarsomeres 5 brownish, otherwise legs yellow. Hind trochanter lacking spinules. Fore femur markedly thickened, with rows of moderately long antero- and posteroventral bristles. Fore tibia spindle-like, with ordinary setation. Mid femur slender, with 2 strong, moderately long bristles near middle and 1 long, subapical bristle anteriorly. Mid tibia with black ventral spinules, lacking prominent bristles. Hind femur (viewed laterally) evenly thickened towards middle, with 1 row of short anteroventral bristles (4-5 subapicals longer) and some dorsal erect bristles near base. Hind tibia with unmodified posterior apical comb, clothed in ordinary setulae. Fore and mid tarsi slender, hind tarsomere 1 thickened. WING. Normally developed, covered with uniform microtrichia; distinctly brownish infuscate. Costal vein with short setulae along anterior margin. R 2+3 long, meeting costa beyond middle of wing. Distance between apices of R 2+3 and R 4+5 about 1.5 times shorter than distance between apices of R 1 and R 2+3 . R 4+5 strongly curved towards costa in apical part. M 1+2 evenly bowed. Halter with black knob and pale stem. ABDOMEN. With tergites (except tergite 8) weakly sclerotised, brownish yellow. Tergite 1 produced laterally into projection bearing 3 short bristles and resting in shallow sclerotised excavation on tergite 2. Sternites 2-4 divided. Gland-like structure not prominent. TERMINALIA. Brown ( Figs 229-231 ). Cerci broadly fused, short, left cercus with 3 very long strong terminal bristles. Epandrium completely divided. Left epandrial lamella fused to hypandrium, with 1 long seta apically. Left surstylus ( Fig. 231 ) with upper lobe divided; lower part with markedly developed surstylar comb and additional short internal apophysis bearing 3 short setae, upper part short, slender, digitiform, with 1 subapical seta. Right epandrial lamella ( Fig.230 ) subtriangular, covered with numerous setae of different lengths. Right surstylus ( Fig. 229 ) not prominent. Hypandrium with 2 very long, strong setae apically. Phallus short. Female Mid tibia lacking ventral spinules. Segment 8 elongate, with tergite and sternite separated; sternite 8 with apical part somewhat hinged. Cercus elongate oval, brownish. Otherwise as in male. Material examined Holotype ♂ SINGAPORE : Sime forest , 16 September 2005 , forest, Mal 1 (reg. 25352, leg. P.G. ; in ZRC). Paratypes Figs 229-231. Stilpon singaporensis sp. nov., ♂. 229 . right epandrial lamella, lateral view. 230 . epandrium with cerci, dorsal view. 231 . left surstylus, lateral view. Scale bar= 0.1 mm. SINGAPORE : 3 ♂♂ , 1 ♀ , Sime forest, 24 March 2005 , forest, sweeping (reg. 25026, leg. P.G.); 3 ♂♂ , 1 ♀ , Sime forest, 15 April 2005 , forest, sweeping (reg. 25061, leg. P.G.); 1 ♂ , 2 ♀♀ , Sime forest, 1 July 2005 , forest, sweeping (reg. 25191, leg. P.G.); 1 ♂ , 2 ♀♀ , Sime forest, 22 July 2005 , forest, Mal 1 (reg. 25270, leg. P.G.); 1 ♂ , Sime forest, 19 August 2005 , forest, Mal 1 (reg. 25299, leg. P.G.); 1 ♂ , Sime forest, 7 September 2005 , forest, Mal 1 (reg. 25334, leg. P.G.); 1 ♂ , 1 ♀ , Sime forest, 18 November 2005 , forest, Mal 1 (reg. 25420, leg. P.G.); 1 ♀ , Sime forest, 3 December 2005 , forest, Mal 1 (reg. 25434, leg. P.G.). Etymology The new species is named after the country of origin, Singapore . Distribution Singapore . Habitat and seasonal occurrence This species was collected in forests, with records from March-April, July-September and November- December. Remarks In having modified male abdominal tergites 1 and 2, the new species is closely related to S. paradoxus Shamshev and Grootaert described from Thailand ( Shamshev & Grootaert 2004b ). However, S. paradoxus can be readily distinguished from S. singaporensis sp. nov. in the slightly arcuate R 4+5 (vs. vein R 4+5 strongly curved towards costa in the new species). Additionally, in the male of S. paradoxus the hind femur is constricted and bent near the middle and the hind tibia bears a long spur-like posterior apical comb. The relationships of both species with Stilpon are not clear.