Wrong side of the leaf: assigning some Lithocolletinae species (Lepidoptera: Gracillariidae) to their proper genera Author Eiseman, Charles S. Author Davis, Donald R. text Zootaxa 2020 2020-03-17 4751 2 201 237 journal article 10.11646/zootaxa.4751.2.1 6d5c6a73-5675-46cf-98a4-ece6ddc8e121 1175-5326 3712951 7692DE47-FE0C-47CA-BF74-10302592AC5F Phyllonorycter holodisci ( Braun, 1939 ) ( Figs. 35–37 ) Lithocolletis holodisci Braun 1939: 282 . Phyllonorycter holodisci (Braun) — Davis 1983: 10 . Leaf mine. An upper-surface blotch, becoming wrinkled when mature ( Figs. 35–36 ). This species has been occasionally observed to form lower-surface mines in California . Host. Rosaceae : Holodiscus discolor (Pursh) Maxim. Distribution. Canada : BC; USA : CA, ID, OR, WA. Review. The type specimen ( Fig. 37 ) was reared from an upper-surface mine on Holodiscus discolor collected in California (Del Norte Co.) on 23 August 1936 . Braun (1939) also collected mines in Washington (Clallam Co.) and Oregon (Lane Co.) but found them to be heavily parasitized and reared no further specimens. De Benedictis et al . (1990) collected mines in California (San Mateo Co.) in April and July but failed to rear adults. They implied that this species forms lower-surface mines, as was later stated by Gates et al . (2002) . A collection of preserved leaf mines from California provided by D.L. Wagner consists primarily of upper-surface mines, but it does indeed include a few underside tentiform mines like those of other Phyllonorycter species (e.g., from Russell Tree Farm, Contra Costa Co., 6 July 1981 ). D.L. Wagner (pers. comm.) has reared adults from both mine types and believes they are conspecific. Mines found in Idaho (Kootenai Co., 27 September 2012 , photographed by CSE; Figs. 35–36 ), Washington (Whatcom Co., 28 July 2016 ; Palmer 2016 ), and British Columbia (Vancouver Island, 10 August 2019 ; Mews 2019 ) were all on the upper leaf surface as described by Braun.