Taxonomic Re-Evaluation Of Sarika Diadema (Dall, 1897) And S. Asamurai (Panha, 1997), Two Endemic Land Snails From Thailand (Pulmonata: Ariophantidae: Macrochlamydinae) Author Sutcharit, Chirasak Author Panha, Somsak text Raffles Bulletin of Zoology 2008 2008-02-29 56 1 95 100 journal article 10.5281/zenodo.5341400 2345-7600 5341400 Sarika asamurai (Panha, 1997) ( Figs. 2C, D ; 3E–I ; 4D–F ) Macrochlamys asamurai Panha, 1996 [1997]: 101–105, Figs. 2a, 2b 1 ( Takun Village , Suratthani , Thailand ; Holotype CUMZ 6325 ). Type material. – Takun Village, Suratthani: Holotype CUMZ 6325 ; Paratype : CUMZ 6326 , 6327 ; FMNH 285043 ; USNM 892000 Other material examined. – Klong Saeng, Suratthani: CUMZ 2584, 2589, 2593, 3617; Wat Tam Wararam, Suratthani: CUMZ 2586, 2590, 2591, 2594, 3618, 4640, 3641; Wat Tam Sok Panthurat, Suratthani: CUMZ 3623, 3624, 3625, 3626; Klong Phanom National Park, Suratthani: CUMZ 3644. Shell. – Shell ( Fig. 2C ) similar to that of S. diadema but slightly larger; transverse sculpture on the upper periphery more dense; descending whorls slightly elevated from suture without shoulder. Genitalia. – Atrium (at) somewhat short. Penis (p) cylindrical, distally thickened from site of penial verge, proximally enclosed in thin penial sheath (ps). Ephiphallic caecum (ec) short, straight and located about half way along the length of the ephiphallic complex. Penial retractor muscle (prm) short, thick and attached at the tip of ephiphallic caecum. Epiphallus (e) small, and slender tube, about twice length of penis; flagellum (fl) short. Vas deference (vd) a small tube running from the spermoviduct and entering at the epiphallus/flagellum junction. ( Fig. 3F ). Internal sculpture of penis similar to that in S. diadema , the most notable difference being a distinctly longer penial pilaster (pp) (about half of penial chamber) in S. asamurai ( Fig. 3G ). The female genital tracts are generally similar to those of S. diadema but with the significant difference that the dart apparatus (da) is located proximally on the vagina at the genital atrium and is covered in black pigmentation ( Fig. 3F ). Allospermatophores (n = 10); only one short spine present at the sperm sac and the tail filament junction( Fig. 3H ). Five specimens contained spermatophores, each with two in the gametolytic sac. Fig. 4. Radula morphology. Sarika diadema from Botanic Garden, Trang: A, central tooth with first to third lateral teeth; B, lateral teeth with bicuspid marginal teeth transition; C, outermost marginal teeth. Sarika asamurai from Klong Phanom National Park, Suratthani: D, central tooth with first and second lateral teeth; E, lateral teeth with bicuspid marginal teeth transition; F, outermost marginal teeth. Numbers indicated order of lateral and marginal teeth. Central tooth indicated by ‘C’. Radula and jaw. – Each row contained about 125 (65-(17- 16)-1-(15-16)-59) teeth. Central and lateral teeth with triangular tricuspid ( Fig. 4D ). Bicuspid marginal teeth from 15–17) ( Fig. 4E ). Radula teeth morphology very similar to that described for S. diadema ; the jaw exhibits significantly less curvature ( Figs. 3I and 4F ). External features. – Head/foot similar in form but markedly different in colour to that of S. diadema . Anterior body black; foot sole reddish and tripartite; mantle collar and thin mantle flaps reddish to dark red. Posterior of body usually entirely reddish or with blackish caudal foss and caudal horn ( Fig. 2D ). Only the anterior and/or caudal end black pigmentation is retained in ethanol preserved specimens. Distribution. – Sarika asamurai has a narrower distribution than S. diadema ( Fig. 1 ). Current records are restricted to the following limestone karst area of Suratthani Province: Klong Phanom and Khao Sok National Parks; Ban Takun village ( type locality); Wat Tam Sok Panthurat; Wat Tam Wararam.