The Drosophila (Sophophora) obscura species group in the Americas (Diptera: Drosophilidae): review, revisions, and three new species Author Grimaldi, David A. text American Museum Novitates 2024 2024-05-16 2024 4015 1 44 https://bioone.org/journals/american-museum-novitates/volume-2024/issue-4015/4015.1/The-Drosophila-Sophophora-obscura-species-group-in-the-Americas-Diptera/10.1206/4015.1.full journal article 10.1206/4015.1 0003-0082 12171426 Drosophila ( Sophophora ) tolteca Patterson and Mainland Figures 4C , 7G , 9L , 12E , 15C Drosophila tolteca Patterson and Mainland , in Mainland and Patterson, 1944: 32 . DIAGNOSIS: Recognizable for the large sex comb on ta1, with generally 7–8 teeth (range of 5–9: Sulerud and Miller, 1966 ); ta2 with one very small, thin “tooth” barely distinguishable from adjacent setae; male ta1 short, length ~0.80× length of ta2. Male genitalia with base of inner ventral epandrial lobe with furrows and dense microtrichia. Female: apex of oviscapt blunt, ovisensilla all short pegs; spermatheca squat, width 2× the height, with apical indentation. TYPE : Types not found. Mainland and Patterson (1944) did not specify a type specimen, merely stating (p. 33) that “ The stock upon which the description is based originated from specimens collected near Cordoba [ Veracruz ].” SPECIMENS EXAMINED: All in AMNH : Mexico: Desietrode de los Leones, Distrito Federal, VII/29/42, G.B. Mainland 1342.1, 1♂ , 1♀ . EL SALVADOR : San Salvador / Jan. 21, 1954 , W.B. Heed , 1♂ ; Volcan Santa Ana , 5670 ft. , 20.7a / Nov. 1953 , W.B. Heed , 1♂ ; Volcan Boqueron , 4500 feet , 91.16 / Feb 25, 1954 , W.B. Heed , 1♂ . NICARAGUA : 11 klm N Matagalpa 57.22 / Santa Maria de Ostuma / June 1954 , W.B. Heed , 1♂ . From culture 14012-0210.0, DNA seq publ by V. Schawaroch , 2002, 5♂♂ ( ASG 23 ), 1♀ ( ASG 24 ) . DISTRIBUTION: Reported from Colombia (near Bogotá : Hunter, 1964 , 1966 ; Villamizar and Alvarez, 2010 ), Ecuador ( Cotopaxi and Pinchincha: Acurio and Rafael, 2009 ), El Salvador ( Heed, 1957 ), and Mexico ( Mainland and Patterson, 1944 ). It is also reported from Coroico (near La Paz) Bolivia , based on a strain studied by Barrio et al. (1992) . I am unaware of any material from Bolivia . The Coroico strain was the only one of D. tolteca studied by Barrio et al. (1992) , so comparisons with Central American strains and specimens have not been made, genetically or morphologically, and the identity of the Bolivia material has not been verified.