On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)
Author
Assing, Volker
text
Beiträge Zur Entomologie = Contributions to Entomology
2016
2016-12-20
66
1
13
111
https://www.contributions-to-entomology.org/article/view/1904
journal article
2419
10.21248/contrib.entomol.66.1.13-111
d2933fad-1cac-4a5a-a558-2633f8314541
0005-805X
6421171
Orphnebius schuelkei
ASSING, 2006
Material examined
:
China
:
1 ♂
,
2 ♀♀
,
Sichuan
,
Emei Shan
,
29°34'N
,
103°21'E
,
1800–2400 m
, sifted,
27.VI.–5.VII.2009
, leg.
Grebennikov
;
1 ♂
,
1 ♀
,
Sichuan
,
Emei Shan
,
29°34'N
,
103°21'E
,
1830 m
, sifted,
26.V.2011
, leg.
Grebennikov
(material in
CAS
, cSme, cAss).
Figs 1–26
. Antenna (
1–21
) and forebody (
22–26
) of
Orphnebius
spp.
of the
O. hauseri
group:
spinans
(
1
,
23
),
cernens
(
2
,
24
),
lunatus
(
3
,
25
),
bakeri
(
4
,
26
),
breviceps
(
5
),
extensus
(
6
),
serratus
(
7
),
integer
(
8
),
bicuspis
, Laos (
9
),
bicuspis
, NE
India (
10
),
fusicollis
(
11
),
retunsus
(
12
),
fodens
(
13
),
opticus
(
14
),
dilatatus
(
15
),
fuscapicalis
(
16
),
grandicollis
(
17
,
22
),
reductus
(
18
),
biformis
, male (
19
),
biformis
, female (
20
),
effeminatus
(
21
). Scale bars: 22–26: 1.0 mm; 1–21: 0.5 mm.
Figs 27–39
. Forebody (
27–34
), abdomen (
35–37
), mesotibia (
38
), and protibia (
39
) of
Orphnebius
spp.
of the
O. hauseri
group:
fuscapicalis
(
27
,
35
),
dilatatus
(
28
,
36
),
latitibialis
(
29
,
38
),
breviceps
(
30
),
extensus
(
31
),
serratus
(
32
,
39
),
integer
(
33
,
37
),
bicuspis
, Laos (
34
). Scale bars: 27–37: 1.0 mm; 38–49: 0.5 mm.
Figs 40–53
. Forebody (
40–50
), abdomen (
51
), and antenna (
52–53
)
of
Orphnebius
spp.
of the
O. hauseri
group (
40–49
) and of the subgenus
Deroleptus
(
50–53
):
opticus
(
40
),
effeminatus
(
41
),
biformis
, male (
42
),
biformis
, female (
43
),
bicuspis
, NE
India (
44
),
retunsus
(
45
),
fusicollis
(
46–47
),
fodens
(
48
),
reductus
(
49
),
niger
(
50–52
),
siamensis
(
53
). Scale bars: 1.0 mm.
Figs 54–83
. Antenna (
54–69
), forebody (
70–73
), abdomen (
74–76
), abdominal segments III–V (
77
,
81
), lateral portion of abdominal segments III–V (
78
,
82
), tergites VI–VII (
79
,
83
), and hind leg (
80
) of
Orphnebius
spp.
of the subgenus
Deroleptus
:
discrepans
(
54
,
73
,
75
),
triapicalis
(
55
,
71
,
76
),
gracilior
(
56
,
72
,
77–80
),
sexcarinatus
(
57
,
81–83
),
spoliatus
(
58
),
cultellatus
(
59
),
septemcuspis
(
60
),
vates
(
61
),
tortus
(
62
),
baccillatus
(
63
),
carinatus
(
64
),
dispar
(
65
),
laticeps
(
66
),
biimpressus
(
67
),
ulcerosus
(
68
),
falagrioides
(
69
),
siamensis
(
70
,
74
). Scale bars: 70–83: 1.0 mm; 54–69: 0.5 mm.
Figs 84–101
. Forebody (
84–90
), abdomen (
91–94
,
97–98
), lateral portion of abdominal segments III–V/VI (
95
,
99
,
101
), and tergite VII (
96
,
100
) of
Orphnebius
spp.
of the subgenus
Deroleptus
:
sexcarinatus
(
84
),
spoliatus
(
85
,
91
),
cultellatus
(
86
,
92–94
),
septemcuspis
(
87
,
95–96
),
tortus
(
88
,
98–100
),
vates
(
89
,
97
),
baccillatus
(
90
,
101
). Scale bars: 84–95, 97–99, 101: 1.0 mm; 96, 100:
0.5 mm
.
Figs 102–118
. Forebody (102–108), abdomen (
109–110
,
112–113
,
115
), lateral portion of abdominal segments III–V/ VII (
111
,
117–118
), and tergites VII(–VIII) (
114
,
116
) of
Orphnebius
spp.
of the subgenus
Deroleptus
:
carinatus
(
102
,
117
),
laticeps
(
103
,
112
),
biimpressus
(
104
,
108
,
113–114
),
ulcerosus
(
105
,
115–116
),
dispar
(
106
,
110–111
),
falagrioides
(
107
,
118
),
baccillatus
(
109
). Scale bars: 102–110, 112–113, 115, 117–118: 1.0 mm; 111, 114, 116:
0.5 mm
.
Figs 119–143
.
Drusilla bifida
(
119–122
),
Rabdotodrusilla pectinata
(
123–125
),
Amaurodera arunica
(
126–127
,
132
),
A. gilvios
(
128
,
133
),
A. reticulata
(
129
,
134
),
A. dentata
(
130
,
135
),
A. parvoculata
(
131
,
136
),
A. thailandensis
(
137–138
),
A. fasciata
(
139
),
A. meorum
(
140
),
A. disparicollis
(
141
),
A. spinans
(
142
), and
A. varicollis
(
143
): forebody (
119
,
123
,
126
,
128–131
); head and pronotum (
120
); antenna (
121
,
124
,
132–143
); abdomen (
122
,
125
,
127
). Scale bars: 1.0 mm.
Figs 144–159
.
Amaurodera thailandensis
(
144–145
),
A. fasciata
(
146
),
A. meorum
(
147
),
A. spinans
(
148
),
A. varicollis
(
149
),
A. disparicollis
, male (
150
),
A. disparicollis
, female (
151
),
Tetrabothrus nilgiricus
(
152–153
),
T. pubescens
,
syntype
(
154–156
),
T. collucatus
(
157–158
), and
T. neoguineensis
,
holotype
(
159
): forebody (
144–152
,
154
,
157
,
159
); abdomen (
153
,
156
); antenna (
155
,
158
). Scale bars: 144–154, 156–157, 159: 1.0 mm; 155, 158:
0.5 mm
.
Figs 160–180
.
Tetrabothrus borneensis
, holotype (
160–162
),
T. punctiventris
(
163–166
),
T. sulawesicus
(
167–169
),
Zyras illecebrosus
, paratype (
170–172
),
Z. gibbus
, holotype (
173–177
), and
Z. quasar
, holotype (
178–180
): forebody (
160
,
163
,
167
,
170
,
173
,
178
); antenna (
161
,
164
,
168
,
171
,
174
,
179
); abdomen (
162
,
165
,
169
,
172
,
175
,
180
); tergite VIII (
166
); tergite III (
176
); tergites VII– VIII (
177
). Scale bars: 160–165, 167–175, 178–180: 1.0 mm; 166, 176–177: 0.5 mm.
Figs 181–196
.
Zyras porrectus
(
181–183
),
Z.wunderlei
(
184–186
),
Pedinopleurus notabilis
(
187–189
), and
Aenictoides derivatus
(
190– 196
): forebody (
181
,
184
,
187
,
190
); antenna (
182
,
185
,
188
,
192
); abdomen (
183
,
186
,
194
); tergites IX–X (
189
); forebody in lateral view (
191
); thorax in ventral view (
193
); abdomen in lateral view (
195
); abdomen in ventral view (
196
). Scale bars: 181–188, 190–196: 1.0 mm; 189: 0.5 mm.
Figs 197–215
.
Orphnebius spinans
(
197–201
),
O. cernens
(
202–206
),
O. lunatus
(
207–210
), and
O. grandicollis
(
211–215
): male segments IX–X (
197
); median lobe of aedeagus in lateral and in ventral view (
198–199
,
202–203
,
208–209
,
212
); ventral process of aedeagus in ventral view (
200
,
214
); paramere (
201
,
204
,
210
,
215
); apex of paramere (
205
); spermatheca (
206
); tergite VIII (
207
,
211
); ventral process of aedeagus in lateral view (
213
). Scale bars:
0.2 mm
.
Figs 216–235
.
Orphnebius fuscapicalis
(
216–219
),
O. nigrapicalis
(
220–222
),
O. bakeri
(
223–225
),
O. breviceps
(
226–228
),
O. dilatatus
(
229–232
), and
O. extensus
(
233–235
): median lobe of aedeagus in lateral and in ventral view (
216–217
,
220–221
,
223
,
226–227
,
229–230
,
233–234
); paramere (
218
,
222
,
224
,
228
,
231
,
235
); apex of paramere (
219
); spermatheca (
225
,
232
). Scale bars:
0.2 mm
.
Figs 236–257
.
Orphnebius serratus
(
236–243
),
O. integer
(
244–250
),
O. latitibialis
(
251–252
), and
O. bicuspis
from Laos (
253– 257
): median lobe of aedeagus in lateral and in ventral view (
236–237
,
244–245
,
253–254
); apex of median lobe in ventral view (
238
,
246
,
255
); paramere (
239
,
247
,
256
); tergite VIII (
240
,
249
,
252
); spermatheca (
241–243
,
251
,
257
); apex of paramerite (
248
); postero-median portion of tergite VIII (
250
). Scale bars: 0.2 mm.
Figs 258–276
.
Orphnebius bicuspis
(
258–260; 258–259
:
India
),
O. opticus
,
syntype
(
261–262
),
O. fusicollis
(
263–268
),
O. reductus
(
269–273
), and
O. biformis
(
274–276
): median lobe of aedeagus in lateral and in ventral view (
258–259
,
263–264
,
269–270
); tergite VIII (
260
,
261
,
266
,
272
,
275
); spermatheca (
262
,
268
,
273
,
276
); paramere (
265
,
271
,
274
); anteromedian portion of sternite VIII (
267
). Scale bars:
0.2 mm
.
Figs 277–295
.
Orphnebius biformis
(
277–278
),
O. retunsus
(
279–282
),
O. fodens
(
283–286
),
O. effeminatus
(
287–292
), and
O. multimpressus
(
293–295
): median lobe of aedeagus in lateral and in ventral view (
277–280
,
283–284
,
287–288
,
293–294
); paramere (
281
,
285
,
289
,
295
); tergite VIII (
282
,
286
,
290
); median portion of tergite VIII (
291
); male tergites IX-X (
292
). Scale bars:
0.2 mm
.
Figs 296–316
.
Orphnebius niger
(
296–298
),
O. siamensis
(
299–303
),
O. discrepans
(
304–306
),
O. triapicalis
(
307–310
),
O. gracilior
(
311–313
),
and
O. sexcarinatus
(
314–316
): tergite VIII (
296
,
302
,
310
,
311
,
314
); female sternite VIII (
297
,
312
,
315
); spermatheca (
298
,
313
,
316
); median lobe of aedeagus in lateral and in ventral view (
299–300
,
304–305
,
307–308
); paramere (
301
,
306
,
309
); male sternite VIII (
303
). Scale bars:
0.2 mm
.
Figs 317–339
.
Orphnebius spoliatus
(
317–324
),
O. cultellatus
(
325–334
: 331, 334:
Laos
), and
O. septemcuspis
(
335–339
): median lobe of aedeagus in lateral and in ventral view (
317–318
,
325–326
,
335–336
); paramere (
319
,
327
,
337
); tergite VIII (
320
,
328
,
338
); male sternite VIII (
321
,
329
,
339
); female sternite VIII (
322
,
330–331
); female segments IX–X in ventral view (
323
,
332
); spermatheca (
324
,
333–334
). Scale bars:
0.2 mm
.
Figs 340–358
.
Orphnebius vates
(
340–344
),
O. tortus
(
345–347
),
O. baccillatus
(
348–350
),
O. carinatus
(
351–353
), and
O. dispar
(
354–358
): median lobe of aedeagus in lateral and in ventral view (
340–341
,
354–355
); paramere (
342
,
356
); tergite VIII (
343
,
345
,
348
,
351
,
357
); male sternite VIII (
344
); female sternite VIII (
346
,
349
,
352
); spermatheca (
347
,
350
,
353
,
358
). Scale bars:
0.2 mm
.
Figs 359–378
.
Orphnebius laticeps
(
359–361
),
O. biimpressus
(
362–364
),
O. ulcerosus
(
365–369
),
O. falagrioides
(
370–372
), and
Drusilla bifida
(
373–378
): tergite VIII (
359
,
362
,
368
,
370
,
374
); female sternite VIII (
360
,
363
,
371
); spermatheca (
361
,
364
,
372
); median lobe of aedeagus in lateral and in ventral view (
365–366
,
376–377
); paramere (
367
); male sternite VIII (
369
,
375
); posteromedian portion or head (
373
); apex of ventral process of aedeagus (
378
). Scale bars: 359–372, 374–377:
0.2 mm
; 373, 378:
0.1 mm
.
Figs 379–395
.
Drusilla bifida
(
379
),
Rabdotodrusilla pectinata
(
380–383
),
Amaurodera arunica
(
384–388
),
A. gilvios
(
389–390
,
395
), and
A. reticulata
(
391–394
): sutural portion of left elytron (
379
); median lobe of aedeagus in lateral and in ventral view (
380– 381
,
385–386
,
389–392
); male tergite VIII (
382
); male sternite VIII (
383
); male pronotum (
384
); spermatheca (
387–388
,
395
); subapical portion of median lobe of aedeagus in dry preparation (
393
); subapical portion of median lobe of aedeagus in transparent light (
394
). Scale bars: 384: 0.5 mm; 379–383, 385–392, 395: 0.2 mm; 393–394: 0.1 mm.
Figs 396–419
.
Amaurodera dentata
(
396–399
),
A. parvoculata
(
400–404
),
A. thailandensis
(
405–410
),
A. fasciata
(
411–414
), and
A. meorum
(
415–419
: 415–416, 419: Khao Yai Nat.Park): median lobe of aedeagus in lateral and in ventral view (
396–397
,
400–402
,
405–409
,
411–412
,
415–418
); spermatheca (
398–399
,
403–404
,
410
,
413–414
,
419
). Scale bars:
0.2 mm
.
Figs 420–441
.
Amaurodera meorum
(
420–422
),
A. disparicollis
(
423–429
),
A. spinans
(
430–431
),
A. varicollis
(
432–434
),
Tetrabothrus nilgiricus
(
435–436
),
T. indicus
(
437–439; 439
:
lectotype
), and
T. inflexus
from
Laos
(
440–441
): spermatheca (
420–422
,
429
,
434
); male pronotum (
423–424
); female pronotum (
425–426
); median lobe of aedeagus in lateral and in ventral view (
427–428
,
430–433
,
435–437
,
440
); ventral process of aedeagus (
438–439
,
441
). Scale bars: 423–426: 1.0 mm; 420–422, 427–441:
0.2 mm
.
Figs 442–458
.
Tetrabothrus collucatus
(
442–443
),
T. borneensis
(
444–449
),
T. punctiventris
(
450–451
),
T. sulawesicus
(
452–454
),
Zyras illecebrosus
,
paratype
(
455–456
), and
Z. quasar
,
holotype
(
457–458
): median lobe of aedeagus in lateral and in ventral view (
442–445
,
450–453
,
455–458
); apico-ventral portion of median lobe in lateral view (
446–448
); male sternite VIII (
449
,
454
). Scale bars:
0.2 mm
.
Figs 459–471
.
Zyras porrectus
(
459–460
),
Z. wunderlei
(
461–465
),
and
Aenictoides derivata
(
466–471
): median lobe of aedeagus in lateral and in ventral view (
459–462
,
468–469
); paramere (
463
,
470
); male tergite VIII (
464
); male sternite VIII (
465
); maxilla (
466
); labium (
467
); apical lobe of paramere (
471
). Scale bars: 459–466, 468–470: 0.2 mm; 467, 471: 0.1 mm.
Comment
: The above specimens represent the first records since the original description, which is based on a male from the Daba
Shan
at the border between
Shaanxi and Chongqing
and one from the Jiajin
Shan
in West
Sichuan
(
ASSING 2006c
). As was to be expected, the previously unknown female primary and secondary sexual characters are similar to those of other species of the
O. hauseri
subgroup.