A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea
Author
Harzhauser, Mathias
Author
Landau, Bernard
text
Zootaxa
2016
4210
1
1
178
journal article
37280
10.11646/zootaxa.4210.1.1
e782e07d-76b7-4e9b-ba34-ed3286254ec6
1175-5326
252966
D39416B8-CF85-440B-84C2-D4380BECC4E3
Kalloconus ponderoaustriacus
(Sacco, 1893)
Figs 3
P, 11G1–G3, 11H1–H3
Conus
(
Rhizoconus
)
ponderosus
Brocc. Var.
III—Hoernes &
Auinger 1879
: 39
(partim), pl. 5, fig. 6 [non
Lautoconus ponderosus
(
Brocchi, 1814
)
].
[
Conus
(
Chelyconus) conoponderosus
Sacc.
]
var.
ponderoaustriaca
Sacc.
—
Sacco 1893b
: 75
[nov. nom. pro
Conus ponderosus
in
Hoernes & Auinger 1879
, pl. 5, fig. 6].
?
Conus ponderosus
Brocc.
var.
Steinabrunnensis
Sacco—Friedberg 1911: 58, text-fig. 12 (non pl. 3, fig. 3) [non
Lautoconus steinabrunnensis
(Sacco, 1893)
].
?
Conus
(
Rhizoconus) ponderosus steinabrunnensis
(Sacco)
—
Csepreghy-Meznerics 1956
: plate captions, pl. 11, figs 11–12.
Conus
(
Chelyconus) ponderosus
var.
ponderoaustriaca
(Sacco)
—Kojumdgieva in
Kojumdgieva & Strachimirov 1960
: 214
, pl. 51, fig. 1.
?
Conus
(
Chelyconus) ponderosus
var.
steinabrunnensis
(Sacco)
—Kojumdgieva in
Kojumdgieva & Strachimirov 1960
: 213
, pl. 50, fig. 8 [non
Lautoconus steinabrunnensis
(Sacco, 1893)
].
Conus ponderosus
Brocc.
—
Eremija 1959
: 187
, pl. 1, figs 5-5a [non
Lautoconus ponderosus
(
Brocchi, 1814
)
].
Conus
(
Rhisoconus)
[sic]
ponderosus
ponderosus
Brocchi—Atanacković 1963: 79
, pl. 15, fig. 1 [non
Lautoconus ponderosus
(
Brocchi, 1814
)
].
?
Conus
(
Dendroconus
)
steindachneri
Hoernes et Auinger—Atanacković 1963: 79
, pl. 14, figs 6–6b.
Conus
(
Chelyconus
)
ponderosus ponderoaustriacus
(Sacco, 1893)
—
Hinculov 1968
: 151
, pl. 38, figs 4a–b.
?
Conus
(
Chelyconus
) aff.
ponderoaustriacus
Sacco, 1893
—
Strausz 1966
: 462
, pl. 69, figs 7–8.
Conus
(
Chelyconus
)
ponderosus ponderoaustriacus
(Sacco, 1893)
—
Nicorici & Sagatovici 1973
: 176
, pl. 27, figs 6–7.
?
Conus
(
Chelyconus
)
ponderosus ponederoaustriacus
[sic] (Sacco, 1893)—
Atanacković 1985
: 177
, pl. 39, figs 10–11.
Conus panderosus
[sic]
ponderoaustriacus
(Sacco)
—
Ionesi & Nicorici 1994
: 62
, pl. 5, figs 5, 7.
?
Conus
(
Chelyconus
)
ponderosus
Brocchi—Chira &
Voia 2001
: 156
, pl. 3, figs 1a–b.
Type
material.
Holotype
NHMW
1854
/0035/0041,
Lăpugiu de Sus
(
Romania
); middle
Miocene
,
Badenian
(
Langhian
).
Studied material.
Holotype and 1 spec. NHMW 1854/0035/0041, 1 spec.
NHMW
1854
/0035/0037,
Lăpugiu de Sus
(
Romania
)
;
2 spec.
NHMW
1856
/0050/0109,
Gainfarn
(
Austria
)
;
5 spec.
NHMW
1855
/0045/0358,
Gainfarn
(
Austria
)
;
9 spec.
NHMW
1861
/0033/0005,
Bujtur
(
Romania
)
.
Illustrated material.
Figs 11
G1–G3, 3P:
holotype
, Lăpugiu de Sus (
Romania
), SL:
50.4 mm
, MD:
27.1 mm
,
NHMW
1854/0035/0041;
Figs 11
H1–H3: Gainfarn (
Austria
), SL:
49.6 mm
, MD:
27.5 mm
,
NHMW
1855/0045/ 0 358.
FIGURE 12.
Principal Components Analysis
|
(PCA) |
of shell |
ratios |
of |
Kalloconus moravicus
|
and
Phasmoconus
|
fuscocingulatus
revealing a clear separation. LWA
|
= angle |
of the last |
whorl, |
LW |
= length width ratio, RD |
= relative diameter, |
RSH = relative height of spire. |
Revised description.
Medium-sized shells with low conical to broad coeloconoid spire; early spire whorls nearly flat, later whorls weakly convex with last spire whorl often slightly more convex; spire whorls smooth; suture channelled. Subsutural flexure shallow, weakly curved, moderately asymmetrical. Spire passing via a rounded shoulder into conical to weakly ventricose last whorl; position of maximum diameter close below shoulder. Last whorl faintly constricted; prominent to weak spiral grooves may appear on lower third; siphonal canal short; siphonal fasciole weak and indistinct. Aperture moderately narrow, posteriorly narrowing and only weakly broadening anteriorly. Colour pattern poorly preserved, but seems to consist of densely spaced continuous spirals.
Shell measurements and ratios.
n =
10 adult
specimens: largest specimen: SL:
59.3 mm
, MD:
34.1 mm
, mean SL:
49.8 mm
(σ = 4.6), mean MD:
27.6 mm
(σ = 2.9), spire angle: µ = 115.2° (σ = 8.1°), last whorl angle: µ = 34.6° (σ = 2.5°), LW: µ = 1.8 (σ = 0.1), RD: µ = 0.64 (σ = 0.03), PMD: µ = 0.9 (σ = 0.02), RSH: µ = 0.1 (σ = 0.03).
Discussion.
Differs from
Lautoconus steinabrunnensis
in the rounded shoulder, convex spire whorls, lower spire and the higher position of the maximum diameter. Its aperture is narrower especially in the posterior part.
Paleoenvironment.
Shallow marine environments; at least at Gainfarn, the species was associated with seagrass.
Distribution in Paratethys.
Badenian (middle Miocene):
Vienna
Basin
: Gainfarn (
Austria
) (
Hoernes & Auinger 1879
);
Transylvanian Basin
: Lăpugiu de Sus (
Romania
) (
Hoernes & Auinger 1879
);
Pannonian Basin:
Hidas, Szob (
Hungary
) (Csepreghy-Meznerics 1950, 1956;
Strausz 1966
);
Zârand Basin
: Minişul de Sus (
Romania
) (
Nicorici & Sagatovici 1973
);
Caransebeş-Mehadia Basin:
Valea Satului (
Romania
) (
Hinculov 1968
);
Buzău
Basin
: Valea Muscelului (
Romania
) (
Ionesi & Nicorici 1994
);
Dacian Basin
: Dobruša (
Bulgaria
) (
Kojumdgieva & Strachimirov 1960
);
Banja Luka Basin
: Jazovac (
Bosnia and Herzegovina
) (
Atanacković 1985
);
Karlovac-Glina Basin:
Glina (
Croatia
) (
Eremija 1959
).