Prodromus of a fern flora for Bolivia. XXXV. Dryopteridaceae Author Kessler, Michael Systematic and Evolutionary Botany, University of Zurich, Zollikerstrasse 107, CH- 8008 Zurich, Switzerland Author Moran, Robbin C. New York Botanical Garden, 2900 Southern Blvd., Bronx, New York, 10458 - 5126, USA Author Mickel, John T. New York Botanical Garden, 2900 Southern Blvd., Bronx, New York, 10458 - 5126, USA Author Matos, Fernando B. Universidade Federal do Paraná, Departamento de Botânica, Caixa Postal 19031, 81531 - 980, Curitiba, PR, Brasil Author Smith, Alan R. Systematic and Evolutionary Botany, University of Zurich, Zollikerstrasse 107, CH- 8008 Zurich, Switzerland & University Herbarium, 1001 Valley Life Sciences Bldg. # 2465, University of California, Berkeley, CA 94720 - 2465, USA & Systematic and Evolutionary Botany, University of Zurich, Zollikerstrasse 107, CH- 8008 Zurich, Switzerland text Phytotaxa 2018 2018-06-11 353 1 448 450 http://dx.doi.org/10.11646/phytotaxa.353.1.1 journal article 10.11646/phytotaxa.353.1.1 1179-3163 Polystichum cochleatum (Klotzsch) Hieron., Bot. Jahrb. Syst. 34: 452. 1904 . = Polystichum pflanzii Hieron., Bot. Jahrb. Syst. 43: 179. 1913 (lectotypification needed). = Polystichum pycnolepis (Klotzsch) T.Moore sensu Tryon & Stolze (1991) . Range: —Andes from Venezuela to Bolivia ( CH , CO, LP). Ecology: —Common; terrestrial in open, rocky hillsides, in rock crevices, at the edge of large rocks, and in grassy or shrubby slopes; (1700–) 3000–4600 m . Notes: —Because the pinnule margins are most strongly revolute in their middle, the pinnules appear sharply triangular. There are morphologically intermediate specimens with P. lehmannii , P. nudicaule , and P. orbiculatum , indicating possible hybridization. These putative hybrids resemble their suspected parents but have the large, dark, involute scales and thick pinnule texture of P. cochleatum .