Taxonomic revision and cladistic analysis of ghost spiders of the genus Tafana Simon, 1903 (Araneae: Dionycha, Anyphaenidae), with the descriptions of twelve new species Author De Oliveira, Luiz Fernando M. 06C42876-2925-4007-A19D-053BD0058F25 Laboratório de Coleções Zoológicas, Instituto Butantan - Av. Vital Brazil, 1500, 05503 - 900, São Paulo, SP, Brazil. & Instituto de Biociências, Universidade de São Paulo, Rua do Matão, 321, 05508 - 090, São Paulo, SP, Brazil. luiz.moura.lfm@gmail.com Author Brescovit, Antonio Domingos D5B81D79-AFAE-47B1-8A6E-DAB448A24BCC Laboratório de Coleções Zoológicas, Instituto Butantan - Av. Vital Brazil, 1500, 05503 - 900, São Paulo, SP, Brazil. antonio.brescovit@butantan.gov.br text European Journal of Taxonomy 2021 2021-03-31 742 1 77 http://dx.doi.org/10.5852/ejt.2021.742.1291 journal article 7531 10.5852/ejt.2021.742.1291 62f9dc57-bade-4fbc-8af7-f92ff107593c 2118-9773 4655404 23424EC3-8440-4C72-AD9E-C1EC3087F7CA Tafana pittieri sp. nov. urn:lsid:zoobank.org:act: 5763F405-387A-4B81-B2A2-D2A309390260 Figs 28 , 33F , 35 Diagnosis Males of Tafana pittieri sp. nov. resemble those of the silhavyi group by having the secondary looping in the middle in the tegulum, but differ from them by having the large laminar, ring-shaped apophysis at the base of the embolus and retrolateral tibial apophysis short and truncated ( Fig. 28C–E ). Females differ from the other species by having the lateral lobes wing-shaped in the median region, covering part of the atrium, and secondary spermathecae with slender shape ( Fig. 28F–G , 33F ). Etymology The specific name is a patronym in honor of Henri Pittier, a botanist, geographer and ethnologist who devoted himself to the study of the flora and fauna in the National Park Henri Pittier, where the specimen was collected. In the year of 1953 the park (until then called Rancho Grande) was renamed in honor of H. Pittier. Material examined Holotype VENEZUELA • 1 ♂ ; Aragua , Rancho Grande, current National Park Henri Pittier ; [ 10°20′58″ N , 67°41′08″ W ]; 8–15 Apr. 1945 ; M. Beebe et. al. leg.; AMNH . Paratype VENEZUELA • 1 ♀ ; same collection data as for holotype; AMNH . Description Male ( holotype , AMNH) COLOR. Exemplar probably discolored. Carapace orange with cephalic region dark ( Fig. 28A ). Labium, sternum and endites yellow. Chelicerae reddish. Legs I–IV yellow, darkest at tarsi. Abdomen yellow with spots in dorsal view ( Fig. 28A ). MEASUREMENTS. Total length 6.3, carapace length 2.8, width 2.21. Clypeus height 0.08. Eye diameters and interdistances: AME 0.1, ALE 0.12, PME 0.1, PLE 0.12; AME–AME 0.04, AME–ALE 0.04, PME– PME 0.12, PME–PLE 0.1, ALE–PLE 0.04. Chelicerae 1.1 long, with four promarginal teeth and four retromarginal denticles. LEG MEASUREMENTS. I—femur 3.0/ patella 1.1/ tibia 3.5/ metatarsus 3.12 /tarsus 1.2/ total 11.92; II— 2.7/ 1.1/ 2.4 /2.8 / 0.93 /9.93; III—1.98 / 0.8/ 1.53 / 2.0/ 0.6/ 6.91; IV—2.7/ 0.9 /2.5 / 3.1/ 0.7/ 9.9. LEG SPINATION. I—tibia v2-2-2, p1-1-0, r1-1-0, metatarsus v2-0-0, p1-0-0, r1-0-0; II—tibia v2-2-2, p1-1-0, r1-1-0, metatarsus p1-0-0, r1-0-0; III—tibia p1-1-1, r1-1-1; IV—tibia p1-1-1. PALP. Cymbium with large retrolateral projection, embolus long with large base and narrow apex; median apophysis long and curved apically; ventral tegular process short and hyaline; tegulum without basal protuberance; sperm duct with secondary looping in horizontal position; median apophysis laminar, long and curved at the apex ( Fig. 28C–E ). ABDOMEN. Length 3.4, epigastric furrow 0.5 from tracheal spiracle, spiracle 1.9 from base of spinnerets. Female ( paratype , AMNH) COLOR. Exemplar probably discolored. Coloration as in males of this species, except for chelicerae reddish brown ( Fig. 28B ). MEASUREMENTS. Total length 5.4, carapace length 2.2, width 1.8. Clypeus height 0.06. Eye diameters and interdistances: AME 0.08, ALE 0.1, PME 0.1, PLE 0.1; AME–AME 0.04, AME–ALE 0.02, PME– PME 0.14, PME–PLE 0.08, ALE–PLE 0.04. Chelicerae 2.2 long, with four promarginal teeth and four retromarginal denticles. LEG MEASUREMENTS. I—femur 1.9 /patella 0.8/ tibia 1.8/ metatarsus 1.6/ tarsus 0.8 /total 6.9; II— 1.5/ 0.6/ 1.05 / 1.4/ 0.6 /5.15; III—1.2/ 0.5/ 0.7 /1.1 /0.4 / 3.9; IV—1.7/ 0.7 /1.6 / 2.0/ 0.52 / 6.52. LEG SPINATION. I—tibia v2-2-2, p1-1-0, r1-1-0, metatarsus v2-0-0, p1-0-0, r1-0-0; II—tibia v2-2-2, p0-1-0, r1-1-0, metatarsus p1-0-0, r1-0-1; III—tibia v1-2-2, p1-1-0, r1-1-0, metatarsus v2-1-2, p0-1-1, r0-1-1; IV—tibia v1-2-2, p1-1-1. EPIGYNUM. Triangular hood with sclerotized border; atrium short; epigynal plate not covered by cuticle, covered in the posterior region; lateral lobes sinuous with wing form in the median region. Internally, copulatory ducts short and curved; primary spermathecae oval and separated; secondary spermathecae small in the posterior region of the copulatory ducts; fertilization ducts longer than primary spermathecae ( Figs 28F–G , 33F ). ABDOMEN. Length 3.2, epigastric furrow 0.6 from tracheal spiracle, spiracle 1.1 from base of spinnerets. Distribution Known only from Aragua , Venezuela ( Fig. 33 ).