<document id="A563F2B8FAEB26074FEB99C83FBD01CA" ID-CLB-Dataset="7531" ID-DOI="10.5852/ejt.2021.742.1291" ID-GBIF-Dataset="62f9dc57-bade-4fbc-8af7-f92ff107593c" ID-ISSN="2118-9773" ID-Zenodo-Dep="4655404" ID-ZooBank="23424EC3-8440-4C72-AD9E-C1EC3087F7CA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617262825393" checkinUser="plazi" docAuthor="De Oliveira, Luiz Fernando M. &amp; Brescovit, Antonio Domingos" docDate="2021" docId="03CF87ADFF8E6608FD3F9549FE81FB62" docLanguage="en" docName="ejt-742_1-77.pdf" docOrigin="European Journal of Taxonomy 742" docSource="http://dx.doi.org/10.5852/ejt.2021.742.1291" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tafana silhavyi MNHN" docType="treatment" docVersion="7" lastPageNumber="39" masterDocId="FFF6FFD5FFA8662EFF99965EFF99FFCF" masterDocTitle="Taxonomic revision and cladistic analysis of ghost spiders of the genus Tafana Simon, 1903 (Araneae: Dionycha, Anyphaenidae), with the descriptions of twelve new species" masterLastPageNumber="77" masterPageNumber="1" pageNumber="39" updateTime="1698935806814" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FC8B1E1BB132B8368F09241ECD16CCE9">Taxonomic revision and cladistic analysis of ghost spiders of the genus Tafana Simon, 1903 (Araneae: Dionycha, Anyphaenidae), with the descriptions of twelve new species</mods:title>
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<mods:namePart id="9EE9DC8EE71B7C6A497DAB0B0E901844">De Oliveira, Luiz Fernando M.</mods:namePart>
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<mods:affiliation id="3D64E8A4F2DC78C768C97D3E4AA26FE0">Laboratório de Coleções Zoológicas, Instituto Butantan - Av. Vital Brazil, 1500, 05503 - 900, São Paulo, SP, Brazil. &amp; Instituto de Biociências, Universidade de São Paulo, Rua do Matão, 321, 05508 - 090, São Paulo, SP, Brazil.</mods:affiliation>
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<mods:namePart id="D2C585AC4CDF3C43823AA91BCB2C0197">Brescovit, Antonio Domingos</mods:namePart>
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<paragraph id="8BD936BBFF8E6608FD3F9549FC14FCFE" blockId="38.[678,909,790,817]" box="[678,909,790,817]" pageId="38" pageNumber="39">
<heading id="D09181D7FF8E6608FD3F9549FC14FCFE" box="[678,909,790,817]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="39" reason="2">
<emphasis id="B912EAA9FF8E6608FD3F9549FC14FCFE" bold="true" box="[678,909,790,817]" pageId="38" pageNumber="39">
The 
<taxonomicName id="4C664D38FF8E6608FD479548FCA2FCFE" ID-CoL="54JLW" authorityName="MNHN" baseAuthorityName="Caporiacco" baseAuthorityYear="1955" box="[734,827,790,817]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="silhavyi">
<emphasis id="B912EAA9FF8E6608FD479548FCA2FCFE" bold="true" box="[734,827,790,817]" italics="true" pageId="38" pageNumber="39">silhavyi</emphasis>
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group
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</paragraph>
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<subSubSection id="C37C6530FF8E6608FF249508FEF9FBEF" pageId="38" pageNumber="39" type="diagnosis">
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<heading id="D09181D7FF8E6608FF249508FEAFFCBF" bold="true" box="[189,310,854,880]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="B912EAA9FF8E6608FF249508FEAFFCBF" bold="true" box="[189,310,854,880]" pageId="38" pageNumber="39">Diagnosis</emphasis>
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<paragraph id="8BD936BBFF8E6608FF2495DCFEF9FBEF" blockId="38.[188,1400,854,1056]" pageId="38" pageNumber="39">
Males of this group can be recognized by having the apophysis at the base of the embolus curved and laminar (
<figureCitation id="135D2A3EFF8E6608FEBE95FDFE32FC72" box="[295,427,931,957]" captionStart="Fig" captionStartId="15.[189,232,1752,1778]" captionTargetBox="[318,1270,265,1732]" captionTargetId="figure-157@15.[315,1272,265,1735]" captionTargetPageId="15" captionText="Fig. 9. A, C–D. Tafana silhavyi (Caporiacco, 1955), ♂ (MNHN 11312). A. Left palp unexpanded, ventral view (arrow indicating character). C. Left palp expanded, prolateral view. D. Left palp expanded, retrolateral view (arrow indicating character). B. Tafana riveti (MCZ), ♂, left palp. E–F. Tafana riveti (MCZ), ♂. E. Left palp, retrolateral view. F. Left palp, prolateral view (arrow indicating character). Abbreviations: 1= primary loop of the sperm duct; 2= secondary loop of the sperm duct; ABE=apophysis of base of the embolus; BH =basal haematodocha; C=cymbium; Ch =character; E =embolus; EP=embolic process; F = fundus; MA=median apophysis; MH= median haematodocha; P=petiole; SD = sperm duct; ST =subtegulum; T=tegulum. Scale bars: A–B, E–F =0.25 mm; C–D=0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4655426" httpUri="https://zenodo.org/record/4655426/files/figure.png" pageId="38" pageNumber="39">Figs 9C–D</figureCitation>
, 
<figureCitation id="135D2A3EFF8E6608FE2E95FDFE73FC72" box="[439,490,931,957]" captionStart="Fig" captionStartId="39.[189,232,1752,1778]" captionTargetBox="[189,1398,423,1708]" captionTargetId="figure-175@39.[189,1398,423,1708]" captionTargetPageId="39" captionText="Fig. 22. Tafana silhavyi (Caporiacco, 1955) (A, C–E= ♂, MNHN 11312; B, F–G=♀, MNHN 11312). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view. D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view (arrow indicating character). G. Vulva, dorsal view. Abbreviations: A= atrium; ABE =apophysis of base of the embolus; CD=copulatory ducts; CO =copulatory opening; Ch =character; E= embolus; FD =fertilization ducts; H =hood; LL=lateral lobes; MA =median apophysis; PA=patellar apophysis; RP=retrolateral projection; RTA=retrolateral tibial apophysis; S1=primary spermathecae; S2 =secondary spermathecae; SD =spermduct;T =tegulum; VTP =ventral tegular process. Scale bars: A–B =2.16 mm; C–G =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655469" httpUri="https://zenodo.org/record/4655469/files/figure.png" pageId="38" pageNumber="39">22C</figureCitation>
, 
<figureCitation id="135D2A3EFF8E6608FE6F95FDFDB3FC72" box="[502,554,931,957]" captionStart="Fig" captionStartId="40.[189,232,1890,1916]" captionTargetBox="[189,1398,1429,1855]" captionTargetId="figure-441@40.[189,1398,1429,1855]" captionTargetPageId="40" captionText="Fig. 23. Tafana silhavyi (Caporiacco, 1955), ♂ (MNHN 13312). A. Left palp, ventral view. B. Left palp, frontal view. C. Left palp, retrolateral view. Abbreviations: ABE =apophysis of base of the embolus; C =cymbium; E= embolus; MA=median apophysis; RP=retrolateral projection; RTA=retrolateral tibial apophysis; T =tegulum. Scale bars: A, C =0.4 mm; B=0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4655475" httpUri="https://zenodo.org/record/4655475/files/figure.png" pageId="38" pageNumber="39">23B</figureCitation>
, 
<figureCitation id="135D2A3EFF8E6608FDAF95FDFD16FC71" box="[566,655,931,958]" captionStart="Fig" captionStartId="45.[189,232,1716,1742]" captionTargetBox="[189,1398,817,1677]" captionTargetId="figure-340@45.[189,1398,817,1677]" captionTargetPageId="45" captionText="Fig. 26. Tafana maracay sp. nov. (A–C, F =♂, MNHN 11314; D–E =♀, MNHN 11314). A. Left palp, ventral view (arrow indicating character). B. Left palp, ventral view. C. Left palp, retrolateral view. D. Vulva, dorsal view. E. Vulva, dorsal view, details of spermathecae. F. Chelicerae, ventral view. Abbreviations:ABE =apophysis of base of the embolus; bTP=basal tegular protuberance; C=cymbium; CD =copulatory ducts; CG=cuticular glands; Ch =character; CO =copulatory opening; E =embolus; FD = fertilization ducts; GD =gland ducts; H =hood; MA =median apophysis; RP=retrolateral projection; RTA =retrolateral tibial apophysis; S1= primary spermathecae; S2 =secondary spermathecae; ST = subtegulum; T= tegulum; VTP=ventral tegular process. Scale bars: A, C, F =0.5 mm; B =0.3 mm; D =0.1 mm; E =0.03 mm." figureDoi="http://doi.org/10.5281/zenodo.4655485" httpUri="https://zenodo.org/record/4655485/files/figure.png" pageId="38" pageNumber="39">26A–B</figureCitation>
); the sperm duct forming an inverted ‘S-shape’ in the male palp not expanded, with secondary looping in the middle of the tegulum (
<figureCitation id="135D2A3EFF8E6608FC72959AFBD7FC11" box="[1003,1102,964,990]" captionStart="Fig" captionStartId="15.[189,232,1752,1778]" captionTargetBox="[318,1270,265,1732]" captionTargetId="figure-157@15.[315,1272,265,1735]" captionTargetPageId="15" captionText="Fig. 9. A, C–D. Tafana silhavyi (Caporiacco, 1955), ♂ (MNHN 11312). A. Left palp unexpanded, ventral view (arrow indicating character). C. Left palp expanded, prolateral view. D. Left palp expanded, retrolateral view (arrow indicating character). B. Tafana riveti (MCZ), ♂, left palp. E–F. Tafana riveti (MCZ), ♂. E. Left palp, retrolateral view. F. Left palp, prolateral view (arrow indicating character). Abbreviations: 1= primary loop of the sperm duct; 2= secondary loop of the sperm duct; ABE=apophysis of base of the embolus; BH =basal haematodocha; C=cymbium; Ch =character; E =embolus; EP=embolic process; F = fundus; MA=median apophysis; MH= median haematodocha; P=petiole; SD = sperm duct; ST =subtegulum; T=tegulum. Scale bars: A–B, E–F =0.25 mm; C–D=0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4655426" httpUri="https://zenodo.org/record/4655426/files/figure.png" pageId="38" pageNumber="39">Figs 9A</figureCitation>
, 
<figureCitation id="135D2A3EFF8E6608FBC2959AFB17FC11" box="[1115,1166,964,990]" captionStart="Fig" captionStartId="39.[189,232,1752,1778]" captionTargetBox="[189,1398,423,1708]" captionTargetId="figure-175@39.[189,1398,423,1708]" captionTargetPageId="39" captionText="Fig. 22. Tafana silhavyi (Caporiacco, 1955) (A, C–E= ♂, MNHN 11312; B, F–G=♀, MNHN 11312). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view. D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view (arrow indicating character). G. Vulva, dorsal view. Abbreviations: A= atrium; ABE =apophysis of base of the embolus; CD=copulatory ducts; CO =copulatory opening; Ch =character; E= embolus; FD =fertilization ducts; H =hood; LL=lateral lobes; MA =median apophysis; PA=patellar apophysis; RP=retrolateral projection; RTA=retrolateral tibial apophysis; S1=primary spermathecae; S2 =secondary spermathecae; SD =spermduct;T =tegulum; VTP =ventral tegular process. Scale bars: A–B =2.16 mm; C–G =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655469" httpUri="https://zenodo.org/record/4655469/files/figure.png" pageId="38" pageNumber="39">22C</figureCitation>
, 
<figureCitation id="135D2A3EFF8E6608FB02959AFB49FC10" box="[1179,1232,964,991]" captionStart="Fig" captionStartId="44.[189,232,1824,1850]" captionTargetBox="[189,1398,499,1784]" captionTargetId="figure-180@44.[189,1398,499,1784]" captionTargetPageId="44" captionText="Fig. 25. Tafana maracay sp. nov. (A, C–E = ♂, MNHN 11314; B, F–G= ♀, MNHN 11314). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view (arrow indicating character). D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: CD=copulatory ducts; Ch =character; CO=copulatory opening; FD = fertilization ducts; H =hood; PA = patellar apophysis; RTA=retrolateral tibial apophysis; S1= primary spermathecae; S2 =secondary spermathecae. Scale bars: A =2.16 mm; B= 3.5 mm; C–G =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655481" httpUri="https://zenodo.org/record/4655481/files/figure.png" pageId="38" pageNumber="39">25C</figureCitation>
). The females can be recognized by having the atrium wide and copulatory ducts large in the anterior region (
<figureCitation id="135D2A3EFF8E6608FA9A95BBFAEAFC30" box="[1283,1395,997,1023]" captionStart="Fig" captionStartId="39.[189,232,1752,1778]" captionTargetBox="[189,1398,423,1708]" captionTargetId="figure-175@39.[189,1398,423,1708]" captionTargetPageId="39" captionText="Fig. 22. Tafana silhavyi (Caporiacco, 1955) (A, C–E= ♂, MNHN 11312; B, F–G=♀, MNHN 11312). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view. D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view (arrow indicating character). G. Vulva, dorsal view. Abbreviations: A= atrium; ABE =apophysis of base of the embolus; CD=copulatory ducts; CO =copulatory opening; Ch =character; E= embolus; FD =fertilization ducts; H =hood; LL=lateral lobes; MA =median apophysis; PA=patellar apophysis; RP=retrolateral projection; RTA=retrolateral tibial apophysis; S1=primary spermathecae; S2 =secondary spermathecae; SD =spermduct;T =tegulum; VTP =ventral tegular process. Scale bars: A–B =2.16 mm; C–G =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655469" httpUri="https://zenodo.org/record/4655469/files/figure.png" pageId="38" pageNumber="39">Figs 22G</figureCitation>
, 
<figureCitation id="135D2A3EFF8E6608FF259258FF68FBEF" box="[188,241,1030,1056]" captionStart="Fig" captionStartId="45.[189,232,1716,1742]" captionTargetBox="[189,1398,817,1677]" captionTargetId="figure-340@45.[189,1398,817,1677]" captionTargetPageId="45" captionText="Fig. 26. Tafana maracay sp. nov. (A–C, F =♂, MNHN 11314; D–E =♀, MNHN 11314). A. Left palp, ventral view (arrow indicating character). B. Left palp, ventral view. C. Left palp, retrolateral view. D. Vulva, dorsal view. E. Vulva, dorsal view, details of spermathecae. F. Chelicerae, ventral view. Abbreviations:ABE =apophysis of base of the embolus; bTP=basal tegular protuberance; C=cymbium; CD =copulatory ducts; CG=cuticular glands; Ch =character; CO =copulatory opening; E =embolus; FD = fertilization ducts; GD =gland ducts; H =hood; MA =median apophysis; RP=retrolateral projection; RTA =retrolateral tibial apophysis; S1= primary spermathecae; S2 =secondary spermathecae; ST = subtegulum; T= tegulum; VTP=ventral tegular process. Scale bars: A, C, F =0.5 mm; B =0.3 mm; D =0.1 mm; E =0.03 mm." figureDoi="http://doi.org/10.5281/zenodo.4655485" httpUri="https://zenodo.org/record/4655485/files/figure.png" pageId="38" pageNumber="39">26D</figureCitation>
, 
<figureCitation id="135D2A3EFF8E6608FF649258FECDFBEF" box="[253,340,1030,1056]" captionStart="Fig" captionStartId="57.[187,230,1824,1850]" captionTargetBox="[189,1398,495,1783]" captionTargetId="figure-161@57.[189,1398,495,1783]" captionTargetPageId="57" captionText="Fig. 33. Female genitalia, digested. A. Tafana huatanay sp. nov. (MUSM). B. Tafana ruizi sp. nov. (IBSP 258183). C. Tafana silhavyi (Caporiacco, 1955) (MNHN). D. Tafana maracay sp. nov. (MNHN 11314). E. Tafana arawak sp. nov. (MNHN 11320). F. Tafana pittieri sp. nov. (AMNH). G. Tafana oliviae sp. nov. (MCZ). H. Tafana orinoco sp. nov. (MIZA 07062). Abbreviations: CD =copulatory ducts; FD =fertilization ducts; H =hood; S1 =primary spermathecae; S2 =secondary spermathecae. Scale bars: A–H =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655511" httpUri="https://zenodo.org/record/4655511/files/figure.png" pageId="38" pageNumber="39">33D–F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37C6530FF8E6608FF249218FE81FB62" pageId="38" pageNumber="39" type="discussion">
<paragraph id="8BD936BBFF8E6608FF249218FE10FB90" blockId="38.[189,1397,1093,1197]" box="[189,393,1093,1120]" pageId="38" pageNumber="39">
<heading id="D09181D7FF8E6608FF249218FE10FB90" bold="true" box="[189,393,1093,1120]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="B912EAA9FF8E6608FF249218FE10FB90" bold="true" box="[189,393,1093,1120]" pageId="38" pageNumber="39">Included species</emphasis>
</heading>
</paragraph>
<paragraph id="8BD936BBFF8E6608FF24922CFE81FB62" blockId="38.[189,1397,1093,1197]" pageId="38" pageNumber="39">
<taxonomicName id="4C664D38FF8E6608FF24922CFDCEFB42" authority="(Caporiacco, 1955)" authorityName="MNHN" baseAuthorityName="Caporiacco" baseAuthorityYear="1955" box="[189,599,1138,1165]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="silhavyi">
<emphasis id="B912EAA9FF8E6608FF24922CFEF7FB43" box="[189,366,1138,1164]" italics="true" pageId="38" pageNumber="39">Tafana silhavyi</emphasis>
(
<bibRefCitation id="EFF74B4AFF8E6608FEE2922CFDD4FB42" author="Caporiacco L. Di." box="[379,589,1138,1165]" pageId="38" pageNumber="39" pagination="245 - 448" refId="ref30518" refString="Caporiacco L. Di. 1955. Estudios sobre los aracnidos de Venezuela 2 a parte: Araneae. Acta biologica Venezuelica 1 (16): 245 - 448." type="journal article" year="1955">Caporiacco, 1955</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C664D38FF8E6608FDC6922DFD4BFB43" authority="De Oliveira &amp; Brescovit, 2021" authorityName="De Oliveira &amp; Brescovit" authorityYear="2021" box="[607,722,1138,1164]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="chimire" status="sp. nov.">
<emphasis id="B912EAA9FF8E6608FDC6922DFD4BFB43" box="[607,722,1138,1164]" italics="true" pageId="38" pageNumber="39">T. chimire</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157D2FF8E6608FD40922CFCADFB43" box="[729,820,1138,1164]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C664D38FF8E6608FCA5922DFC27FB43" authority="De Oliveira &amp; Brescovit, 2021" authorityName="De Oliveira &amp; Brescovit" authorityYear="2021" box="[828,958,1139,1164]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="maracay" status="sp. nov.">
<emphasis id="B912EAA9FF8E6608FCA5922DFC27FB43" box="[828,958,1139,1164]" italics="true" pageId="38" pageNumber="39">T. maracay</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157D2FF8E6608FC5C922CFBB9FB43" box="[965,1056,1138,1164]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C664D38FF8E6608FBB1922DFB02FB43" authority="De Oliveira &amp; Brescovit, 2021" authorityName="De Oliveira &amp; Brescovit" authorityYear="2021" box="[1064,1179,1138,1164]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="arawak" status="sp. nov.">
<emphasis id="B912EAA9FF8E6608FBB1922DFB02FB43" box="[1064,1179,1138,1164]" italics="true" pageId="38" pageNumber="39">T. arawak</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157D2FF8E6608FB3B922CFB64FB43" box="[1186,1277,1138,1164]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C664D38FF8E6608FA9C922DFAECFB43" authority="De Oliveira &amp; Brescovit, 2021" authorityName="De Oliveira &amp; Brescovit" authorityYear="2021" box="[1285,1397,1139,1164]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pettieri" status="sp. nov.">
<emphasis id="B912EAA9FF8E6608FA9C922DFAECFB43" box="[1285,1397,1139,1164]" italics="true" pageId="38" pageNumber="39">T. pettieri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157D2FF8E6608FF2492CDFE81FB62" box="[189,280,1171,1197]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>