Taxonomic revision of the monotypic genera of Siculodini (Lepidoptera, Thyridoidea, Thyrididae, Siculodinae) Author Queiroz-Santos, Luziany 0000-0002-3796-2823 Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, P. O. Box 19020, 81.531 - 980, Curitiba, Paraná, Brazil. & luzianyqueiroz @ gmail. com; https: // orcid. org / 0000 - 0002 - 3796 - 2823 luzianyqueiroz@gmail.com Author Casagrande, Mirna Martins 0000-0002-6076-8463 Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, P. O. Box 19020, 81.531 - 980, Curitiba, Paraná, Brazil. & mibras @ ufpr. br; https: // orcid. org / 0000 - 0002 - 6076 - 8463 mibras@ufpr.br Author Mielke, Olaf Hermann Hendrik 0000-0003-3655-4606 Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, P. O. Box 19020, 81.531 - 980, Curitiba, Paraná, Brazil. & omhesp @ ufpr. br; https: // orcid. org / 0000 - 0003 - 3655 - 4606 omhesp@ufpr.br text Zootaxa 2021 2021-01-28 4920 2 235 253 journal article 8391 10.11646/zootaxa.4920.2.5 5676f8ca-c69e-461c-894d-2938f1f5cbbc 1175-5326 4475213 0A5D8BEB-2156-4198-B59E-C99565BCB2E0 Loxiorhiza Warren, 1904 ( Figs 3 , 7 , 11 , 14 , 16 ) Loxiorhiza Warren, 1904 . Novitates Zoologicae, 11 (1): 494; species included: Siculodes cervinalis Pagenstecher, 1892 .— Dalla Torre, 1914 . Lepidopterorum Catalogus 20 : 37.— Gaede, 1936 , in Seitz. Die Gross -Schmetterlinge des Amerikanischen Faunengebietes. Die amerikanischen Spinner und Schwärmer 6 : 1204, fig. 180h (d).— Whalley, 1964 . Annals and Magazine of Natural History, (13) 7 (74): 122.— Whalley & Heppner, 1995 , Thyridoidea, in Becker et al. Checklist: Part 2, Hyblaeoidea–Pyraloidea–Tortricoidea , in Heppner (ed.). Atlas of Neotropical Lepidoptera 3 : 32. Type species : Siculodes cervinalis Pagenstecher, 1892 , by monotypy. Taxonomic history . Warren’s (1904) description of the genus Loxiorhiza included Siculodes cervinalis . He also noted that Zeuzerodes Pagenstecher, 1892 is distinct from Loxiorhiza , justifying the description of the new genus. Diagnosis . Loxiorhiza is distinguished from the remaining genera of Siculodini by: (i) wings with an ocher ground color with irregular dark gray spots, more abundant on the ventral face ( Fig. 3 ); (ii) male with setae on the anal margin; (iii) saccus with setae; (iv) valva with editum; (v) fultura superior sclerotized with spines; (vi) vesica with cornuti; (vii) signum oval and (viii) papillae anales suboval. Redescription . Forewing: length—males 18.6 mm (n = 13), females 18.6 mm (n = 11); male with slightly convex outer margin, female with anterior half slightly concave and posterior half straight, both sexes with diffuse dark gray line between R 5 –M 3 on both sides; retinaculum above Sc. Hindwing: length—males 10.9 mm (n = 13), females 11.3 mm (n = 11); both sexes triangular, nearly half the size of the forewing; dark gray diffused line on both sides between M 1 –M 3 ; dorsal surface with darker distal half; male with a tuft of setae on anal margin close to the base; male with one and female with two bristles in the frenulum. Male genitalia ( Fig. 7 ): tegumen rectangular in dorsal view, with rounded posterior angles, posterior margin with slight medial indentation; saccus suboval in lateral view, with numerous long and thin setae in the distal portion; uncus triangular, base with small lateral lobes and with posterior tip curved ventrally, thin and short setae immediately posterior the base; gnathos thin, U-shaped, membranous in the basal portion, and projected ventrally; valva spatula-shaped, costa slightly concave, rounded apex and ventral margin straight, editum small with thin and long setae, sacculus rectangular and narrow; fultura inferior U-shaped, with rounded ends and short setae; fultura superior sclerotized, with two arms converging posteriorly, forming an inverted V-shape with numerous spines; aedeagus cylindrical and curved in lateral view and, in dorsal view, bilobed distally, with proximal opening for the ejaculatory ductus rounded and vesica with small cornuti. Female genitalia ( Fig. 11 ): tergum VIII rectangular in lateral view, lamella antevaginalis slightly sclerotized, ostium membranous; posterior apophyses longer than the anterior apophyses; bursa copulatrix with the beginning of the ductus bursae sclerotized and the corpus bursae four times smaller than the ductus, signum ( Fig. 14 ) oval, with numerous sclerotized spines; papillae anales suboval, as long as tall, covered with thin and short setae.