The genus Leucophenga (Diptera, Drosophilidae), part VIII: twenty-one species from the Oriental region, with morphological and molecular evidence Author Huang, Jia Author Wang, Yalian Author O’Grady, Patrick Michael Author Su, Yirui Author Chen, Hongwei hongweic@scau.edu.cn text Zootaxa 2018 2018-10-19 4503 1 1 70 http://dx.doi.org/10.11646/zootaxa.4503.1.1 journal article 10.11646/zootaxa.4503.1.1 1175-5326 5297894 5F058E32-AEF1-4070-8CC8-177B4CE140BA Leucophenga helvipecta Huang & Chen , sp. nov. ( Figs. 3H , 7M–O , 11A–C , 15I, 15J , 31 ) Diagnosis. This species is most similar to L . forcipula sp. nov. in having the wing entirely hyaline ( Fig. 3E, H ), the colour patterns of abdominal tergites in females ( Figs. 10H , 11C ), the shape and pubescence of epandrium, cercus, and surstylus ( Figs. 28A , 31A ), and the shape of paramere ( Figs. 28C , 31C ), but can be distinguished from the latter in having the abdominal 3rd tergite not shortened in males ( Fig. 11A, B ); aedeagus laterally with a long process at same its height ( Fig. 31D ). Description. Ocellar triangle brownish, with ca. 2 setae above ocellar setae. Frons brownish anteriorly, brownish yellow posteriorly ( Fig. 7M ). Pedicel dark yellow; first flagellomere yellow ( Fig. 7M ). Clypeus entirely dark yellow. Palpus yellow ( Fig. 7M ). Mesonotum mostly dark yellow ( Fig. 7N ). Postpronotal lobe yellow ( Fig. 7O ). Acrostichal setulae in ca. 12–14 irregular rows ( Fig. 7N ). Scutellum entirely brownish yellow ( Fig. 7N ). Pleura with a brownish yellow to brownish, indistinct longitudinal stripe above ( Fig. 7O ). Katepisternum and mesopleuron mostly yellow ( Fig. 7O ). Wing ( Fig. 3H ): Costal vein between R 2+3 and R 4+5 distally with ca. 4–6 peglike spinules on ventral surface; R 4+5 and M 1 nearly parallel distally; halter usually yellowish to yellow, sometimes with a brownish yellow patch at knob ( Fig. 7N ). Abdominal tergites dark brown to dark brown, 2nd tergite mostly yellow, 3rd to 5th tergites usually yellow submedially and sublaterally, 4th and 5th tergites with yellow patches submedially or sublaterally in males ( Fig. 11A, B ), 4th and 5th tergites mostly brown in females ( Fig. 11C ). Male terminalia: Epandrium with ca. 4 setae near posterior margin per side ( Fig. 31A ). Hypandrium with 1 sensillum ( Fig. 31B ). Paramere slender, narrow basally, with ca. 5 sensilla subdorsally ( Fig. 31C ). Measurements and indices. BL = 2.93 mm in holotype (range in 4♂ and 5♀ paratypes : 2.80–3.07 in ♂ , 2.40– 2.93 in ♀ ), ThL = 1.40 mm ( 1.27–1.47 in ♂ , 1.13–1.40 in ♀ ), WL = 2.37 mm ( 2.33–2.53 in ♂ , 2.03–2.68 in ♀ ), WW = 1.17 mm ( 1.12–1.20 in ♂ , 0.90–1.33 in ♀ ), arb = 7/2–3 (6–8/2–3), avd = 0.97 (0.80–0.93), adf = 2.13 (2.00– 2.31), flw = 1.87 (1.60–2.14), FW/HW = 0.41 (0.38–0.45), ch/o = 0.05 (0.05–0.07), prorb = 0.073 (0.68–0.95), rcorb = 0.68 (0.54–0.68), vb = 0.52 (0.40–0.60), dcl = damaged (0.33–0.44), presctl = 0.58 (0.47–0.67), sctl = 1.16 (1.14–1.25), sterno = 0.78 (0.67–0.83), orbito = 2.22 (2.00–2.25), dcp = 0.23 (0.16–0.26), sctlp = 0.93 (0.86–1.07), C = 2.26 (2.16–2.47), 4c = 1.34 (1.27–1.43), 4v = 2.21 (2.13–2.55), 5x = 1.38 (1.36–1.57), ac = 1.86 (1.80–2.00), M = 0.76 (0.73–0.83), C3F = 0.79 (0.66–0.79). Type specimens. Holotype ♂ ( SCAU , No. 128594), JAPAN : Iriomote Is. , Ryukyu Is. , 24°20'N , 123°49'E , alt. 330m , 16.vi.1999 , ex tree trunk, HW Chen . Paratypes : JAPAN : 4♂ , 3♀ ( SCAU , Nos 128595–601), same data as holotype ; CHINA : 1♀ ( SCAU , No. 128602), Baping , Fusui , Guangxi , 22°29'N , 107°54'E , alt. 110m , 18.viii.2004 , ex tree trunk, HW Chen ; 1♂ ( SCAU , No. 128603), Yixiang , Pu'er , Yunnan , 22°47'N , 101°02'E , alt. 1200m , 17.xi.2011 , ex tussock, JJ Gao ; 1♀ ( SCAU , No. 128604), Hesong , Menghai , Yunnan , 21°50'N , 100°06'E , alt. 1700m , 28.iii.2011 , ex tree trunk, YR Su . Distribution. China ( Guangxi , Yunnan ), Japan (Ryukyu Is.). Etymology. A combination of the Latin words: “ helvus ” (= honey gold) and “ pectus ” (= thorax), referring to the mesonotum mostly dark yellow.