One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations
Author
Zamani, Alireza
A21C0B82-E2D9-4402-9408-D6160843DAF4
Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland. Department of Molecular Biology and Genetics, Democritus University of Thrace, Dragana, 68100 Alexandroupolis, Greece. Perm State University, Bukireva Street 15, Perm, 614600, Russia. Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, 685000, Russia. Department of Zoology & Entomology, University of the Free State, Bloemfontein 9300, South Africa.
zamani.alireza5@gmail.com
Author
Chatzaki, Maria
2FBB48ED-84FE-4386-8F47-C958E9F8A2A1
maria.chatzaki@gmail.com
Author
Esyunin, Sergei L.
2B591857-5FAB-4B7D-B589-6C4977A3A493
Sergei.
Sergei.Esyunin@psu.ru
Author
Marusik, Yuri M.
F215BA2C-5072-4CBF-BA1A-5CCBE1626B08
yurmar@mail.ru
text
European Journal of Taxonomy
2021
2021-06-04
751
1
68
114
http://dx.doi.org/10.5852/ejt.2021.751.1381
journal article
6279
10.5852/ejt.2021.751.1381
4e3af683-d8a6-40de-991b-c4584d66b8cc
2118-9773
4906007
1D27EA41-2396-463A-B1B2-0CCF79C9F3D2
Berinda hoerwegi
sp. nov.
urn:lsid:zoobank.org:act:
C322064D-4C9C-4FE5-A6AE-1403E560CA55
Fig. 2
Diagnosis
Males of this species differ from all congeners by the tibial apophysis being rectangular with a blunt end (vs with tapering tip) and by the distinct form of the median apophysis (= conductor sensu
Panayiotou
et al
. 2010
), with the sclerotized part directed terminally (vs laterally in
B
.
aegilia
Chatzaki, 2002
and
B
.
amabilis
Roewer, 1928
or hook-like in
B
.
ensigera
(O. Pickard-Cambridge, 1874)
,
B
.
cooki
Logunov, 2012
,
B
.
cypria
Chatzaki & Panayiotou, 2010
and
B
.
hakani
Chatzaki & Seyyar, 2010
). Females of
B
.
hoerwegi
sp. nov.
are mostly similar to those of
B
.
aegilia
but can be distinguished by the smaller conical anterior hood (
Ah
) (vs rectangular) and longer copulatory ducts.
Etymology
This species is named after Mr Christoph Hörweg, the curator of Arachnology at NHMW, for helping the first author during his visit to
Vienna
.
Fig. 2.
Berinda hoerwegi
sp. nov.
(A–E = holotype, ♂, MHNG; F–G = paratype, ♂ from Kermanshah Provinve, MHNG; H–L = paratype, ♀ from Ilam Province, MHNG).
A
. Habitus, dorsal view.
B–E
. Palp, ventral, retrolateral and retroventral views.
F
. Apical portion of bulb.
G
. Tibial apophysis, retrolateral view.
H, J–K
. Epigyne, ventral view.
I
. Epigyne, dorsal view.
L
. Same, with schematic course of internal duct system.Abbreviation:
Ah
= anterior hood. Scale bars:A = 1 mm; B–C = 0.2 mm; F–G, K–L = 0.1 mm.
Type material
Holotype
IRAN
•
♂
;
Ilam Province
,
Abdanan
;
32°53′ N
,
47°30′ E
;
Jul. 2015
;
A. Zamani
leg.;
MHNG
.
Paratypes
IRAN
•
1 ♀
; same collection data as for holotype;
MHNG
•
8 ♂♂
;
Kermanshah Province
,
Zarrin Joob Vil.
;
Apr. 2017
;
M. Amiri
leg.;
MHNG
•
1 ♀
;
Kurdistan Province
,
Marivan
;
May 2017
;
A. Zamani
leg.;
MHNG
•
1 ♂
;
Fars Province
,
5 km
N of Persepolis
; 1970;
K. Bilek
leg.;
NHMW
.
Description
Male
(
holotype
) Habitus as in
Fig. 2A
. Total length 4.50. Carapace 1.92 long, 1.50 wide. Eye sizes and interdistance of PMEs: AME = 0.09, ALE = 0.12, PME = 0.12, PLE = 0.12, PME–PME = 0.03. Carapace, chelicerae, sternum, maxillae and labium light reddish brown. Chelicera with 2 pro- and 5 retromarginal teeth. Legs slightly lighter than carapace, without annulations, and with numerous spines (III and IV).Abdomen light brown, without any markings and with a small scutum anteriorly. Spinnerets light gray. Measurements of legs: I: 7.29 (1.77, 1.33, 1.66, 1.50, 1.03), II: 5.35 (1.41, 0.87, 1.13, 1.08, 0.86), III: 4.41 (1.19, 0.68, 0.82, 1.01, 0.71), IV: 7.07 (1.81, 1.06, 1.67, 1.71, 0.82).
Palp as in
Fig. 2B–G
. Femur unmodified; patella slightly longer than tibia (not including apophysis); tibial apophysis as long as tibia, with swollen base and flat anterior part, margins parallel; cymbium 2 times as long as wide, its width as long as tibia; spermophor coiled; median apophysis (= conductor sensu Panayiotou
at al
. 2010) large, sharply pointed; filamentous embolus originates almost from the center of the bulb, makes 360° loop.
Female
(
paratype
from
Ilam Province
)
Carapace 3.05 long, 2.28 wide (abdomen missing). Eye sizes and interdistance of PMEs: AME = 0.13, ALE = 0.15, PME = 0.13, PLE = 0.12, PME–PME = 0.05. Coloration and habitus features as in male. Measurements of legs: I: 8.79 (2.30, 1.73, 1.86, 1.69, 1.21), II: 6.99 (1.92, 1.26, 1.55, 1.34, 0.92), III: 4.41 (1.19, 0.68, 0.82, 1.01, 0.71), IV: 6.04 (1.65, 1.00, 1.12, 1.39, 0.88).
Epigyne as in
Fig. 2H–L
. Epigyne with small, conical anterior hood, and lateral plates with copulatory openings located posteriorly in relation to the hood; copulatory ducts long, extending to the edges of the epigynal plate with two full coils, continuing posteriorly to two small, oval spermathecae.
Distribution
Known from the listed localities in
Ilam
,
Kermanshah
,
Kurdistan
and
Fars
Provinces, western and southcentral
Iran
.