A bee from the middle Miocene Hindon Maar of southern New Zealand (Hymenoptera, Colletidae) Author Engel, Michael S. Division of Invertebrate Zoology, American Museum of Natural History, 200 Central Park West, New York, New York 10024 - 5192 (United States); mengel @ amnh. org mengel@amnh.org Author Kaulfuss, Uwe Georg-August-Universität GÖttingen, Johann-Friedrich-Blumenbach Institut für Zoologie und Anthropologie, Evolution und Biodiversität der Tiere, Untere Karspüle 2, 37073 GÖttingen (Germany) uwe. kaulfuss @ uni-goettingen. de text Zoosystema 2025 2025-01-29 47 3 43 49 https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2025v47a3.pdf journal article 10.5252/zoosystema2025v47a3 1638-9387 276828D7-9B6D-4D40-B381-53583AF6208A Leioproctus ( Otagocolletes ) barrydonovani n. sp. ( Figs 2-4 ) urn:lsid:zoobank.org:act: 8037BA79-7FEA-4BEF-8197-F518CB454571 HOLOTYPE . — New Zealand • ♀ ; OU46558 ; Lacustrine mudstones from Hindon Maar (Langhian; I44/f0392 in the New Zealand Fossil Record File); Dunedin Volcanic Group , 10 km N of Outram , Otago , southern New Zealand ; Department of Geology , University of Otago , Dunedin , New Zealand. TYPE LOCALITY AND HORIZON . — Lacustrine mudstones from Hindon Maar (Langhian; I44/f0392 in the New Zealand Fossil Record File), Dunedin Volcanic Group, 10 km N of Outram, Otago , southern New Zealand . FIG . 3. — Leioproctus ( Otagocolletes ) barrydonovani n. sp. holotype OU46559, habitus of counterpart. Scale bar: 1 mm. ETYMOLOGY . — The specific epithet honours the late Barry J. Donovan (1941-2022), authority on New Zealand’s unique bee fauna (e.g. Donovan 2007 ) and a charming and generous man with whom we had the pleasure of working. This project was just starting with Barry when he passed away ( van Toor et al. 2022 ). He is missed. DIAGNOSIS . — As for the subgenus ( vide supra ). DESCRIPTION Female Body. Total length as preserved (vertex to apex of fragment of tergum II) 6.4 mm ; forewing length as preserved (base to torn apex as apical portion of membrane largely missing including apex of marginal cell) 5.6 mm ; mesoscutum length 2.1 mm , mesoscutellum length 0.4 mm ; metasoma width as preserved 2.7 mm . Integument nearly black as preserved (colouration taphonomically altered but likely dark brown to black in life). Sculpturing not discernible for most sclerites (ental surfaces exposed for many sclerites and thereby not depicting external sculpturing or pubescence), where evident seemingly closely punctate (e.g. small portions of mesoscutum, at least laterally, and metasomal terga but otherwise these sclerites internally exposed). Head. Apparently broad (incomplete and in oblique posterior position so precise dimension impossible to determine but seeming broad based on upper width as preserved); gena narrower than compound eye; malar space apparently about as long as wide (the sclerite at the lower margin of the compound eye is not the base of the mandible and indicates a malar space that is 0.5× as long as wide, or even slightly more; this seems almost certainly the case in the part but is less clear in the counterpart as preserved); mesoscutellum longer than combined lengths of metanotum and basal area of propodeum; metanotum exceedingly short (as preserved), slightly shorter than basal area of propodeum; basal area of propodeum wholly declivitous (best observed on part), almost undifferentiated from posterior surface. FIG . 4. — Leioproctus ( Otagocolletes ) barrydonovani n. sp. holotype OU46559, line drawings of left ( A ) and right ( B ) forewing as preserved. Scale bar: 1 mm. Forewing. All wing veins strong and tubular; marginal cell long (although incomplete as preserved the course of Rs and anterior wing margin demonstrate overall length) ( Figs 2-4 ), longer than pterostigma; prestigma about as long as wide and perhaps about as long as pterostigmal base; pterostigma slender, roughly parallel-sided, not widening apically ( Fig. 4 ), tapering within marginal cell, border within marginal cell distinctly convex ( Fig. 4 ); basal vein (1M) faintly arched, confluent with 1cu-a; 1cu-a straight, oblique, sloping toward wing apex; 1Rs slightly longer than prestigma, oblique relative to Rs+M, much shorter than 1 M; Rs+M nearly straight; first submarginal cell longer than individual lengths of second and third submarginal cells, slightly longer than combined lengths of remaining submarginal cells; prestigmal length of first submarginal cell slightly less than 0.3× pterostigmal length of cell; 1r-rs well distad pterostigmal midlength, near pterostigmal apex, shorter than 3Rs; second submarginal cell slightly narrowed anteriorly, 2Rs longer than 3Rs, 2Rs and 1rs-m not parallel; 1m-cu entering second submarginal cell at cell midlength, basad 1rs-m by 4-5× vein width; 3Rs shorter than 4Rs; anterior border of third submarginal cell longer than anterior border of second submarginal cell; 2rs-m weakly curved distally; posterior border of third submarginal cell about as long as that of second submarginal cell, only slightly longer than anterior border of third submarginal cell; 2m-cu apparently entering third submarginal cell near apex (incomplete but given that majority of posterior border of cell present and not in contact with 2m-cu indicates it must make contact near cell’s apex); 2Cu much longer than 2cu-a. Male Unknown.