Ephemeroptera from the Venezuelan Guayanas´s Uplands: Families Leptophlebiidae, Euthyplociidae and Oligoneuriidae Author Domínguez, Eduardo Instituto de Biodiversidad Neotropical (IBN), CONICET-Universidad Nacional de Tucumán. Miguel Lillo 205. 4.000 Tucumán, Argentina. E-mail: mayfly @ unt. edu. ar Author Grillet, Maria-Eugenia Laboratorio de Biología de Vectores y Parásitos, Instituto de Zoología y Ecología Tropical, Facultad de Ciencias, Universidad Central de Venezuela, AP 47058, Caracas 1041 - A, Venezuela. E-mail: maria. grillet @ ciens. ucv. ve Author Nieto, Carolina Instituto de Biodiversidad Neotropical (IBN), CONICET-Universidad Nacional de Tucumán. Miguel Lillo 205. 4.000 Tucumán, Argentina. E-mail: mayfly @ unt. edu. ar Author Molineri, Carlos Instituto de Biodiversidad Neotropical (IBN), CONICET-Universidad Nacional de Tucumán. Miguel Lillo 205. 4.000 Tucumán, Argentina. E-mail: mayfly @ unt. edu. ar Author Guerrero, Edmundo Museo del Instituto de Zoología Agrícola Francisco Fernández Yépez, Facultad de Agronomia, Universidad Central de Venezuela, Maracay, AP 4579, Maracay, Venezuela. E-mail: edmundo. guerrero @ miza-ucv. org. ve text Zootaxa 2014 2014-07-04 3827 3 301 317 journal article 5314 10.11646/zootaxa.3827.3.1 54637182-8d50-4ef3-ad88-e3ee10f3d4cd 1175-5326 4920666 8C4821D8-072C-453A-ACF6-6EC539AE96AD Paramaka Savage & Domínguez The genus was established for the species Paramaka convexa from Surinam ( Savage & Domínguez, 1992 ). Later, in 2003, Blanco Belmonte et al. described the nymph of the genus. In 2005, Sartori described P. antonni based on nymphs from Guyana . Mariano (2011) described another species, P. pearljam , from nymphs and male imagos from Central-western Brazil . The nymphs of these two species presented a labrum not as expanded, nor curved as in P. convexa , suggesting that this could be a specific character. This is confirmed with the different degree of development presented in the new species described below, plus some allometric variation in this character throughout the nymphal development. Furthermore, the new species described here presents gills only on segments I–VI, without the thin and pointed projection usually present in P. convexa and P. antonii (but see variations in P. convexa below). Another difference between the new species described here and the other species known is the strong development of posterolateral projections on segments VII–IX. This development can be related to the total loss of gills on segment VII.