A new integrated morpho- and molecular systematic classification of Cenozoic radiolarians (Class Polycystinea) - suprageneric taxonomy and logical nomenclatorial acts
Author
Suzuki, Noritoshi
Author
Caulet, Jean-Pierre
Author
Dumitrica, Paulian
text
Geodiversitas
2021
2021-07-08
43
15
405
573
journal article
5275
10.5252/geodiversitas2021v43a15
a8353504-9387-42cf-8d81-8ecacbe9bd90
1638-9395
5101757
urn:lsid:zoobank.org:pub:DC259A19-9B35-4B33-AD9F-44F4E1DA9983
Superfamily
CARPOCANIOIDEA
Haeckel, 1882
n. stat.
Carpocanida
Haeckel, 1882: 427
[below a tribe].
DIAGNOSIS. — Consists of an oval to spindle shape shell with two to three (rarely four) segments. Cephalis small, tending to sink into the thorax.
REMARKS
As no appropriate superfamily name has been proposed for this family, the taxonomic rank of the
Carpocaniidae sensu
De Wever
et al.
(2001)
is used here. However, some partial definitions of this superfamily used byDe
Wever
et al.
(2001)
, “cephalis simple”, are excluded from its definition as the
type
genus of the
Carpocaniidae
is known to have a very complex, cephalic initial spicular system (see remarks for
Carpocaniidae
). The “
Carpocaniidae
” sensu bothPetrushevskaya (1981: 225-226) and
De Wever
et al.
(2001)
used to be a member of the superfamily
Eucyrtidioidea
, but
Matsuzaki
et al.
(2015: 66)
excluded this family from the
Eucyrtidioidea
based on the different cephalic structures observed. This morphological decision was later confirmed by a molecular study (
Sandin
et al.
2019
). The cephalic structure, similar to the
Carpocaniidae
and
Artostrobiidae
, is a complex arch and composed of supplementary rod-systems along the V-rod. As these families are poorly differentiated by molecular studies (99 PhyML bootstrap value 10 000 replicates, BS and 0.68 posterior probabilities), it may not be necessary to separate them at the superfamily level. This superfamily includes the
Carpocaniidae
and probably the
Diacanthocapsidae
in the Cenozoic.