An overview of the Onthophagini from the Arabian Peninsula (Coleoptera: Scarabaeoidea: Scarabaeidae) Author Ziani, Stefano Author Abdel-Dayem, Mahmoud Saleh Author Aldhafer, Hathal Mohammed Author Barbero, Enrico text Zootaxa 2019 2019-08-21 4658 1 1 36 journal article 25961 10.11646/zootaxa.4658.1.1 31d6e081-95c3-4086-a45a-c11555dda3aa 1175-5326 3373040 ADCBCCA2-63C1-42CD-A3C9-C14F954F4A21 Onthophagus ( Indonthophagus ) nitidulus Klug, 1856 [ Fig. 13 ] Literature quotes . SAUDI ARABIA : without locality ( Gestro 1889 ); Makkah Province , Jeddah ( Zunino 1981 ). YEMEN : Lahij Governorate , Lahej [Al Houta]. Al Hudaydah Governorate , Kamaran Island (Scott in Paulian 1948 ). UNITED ARAB EMIRATES : Abu Dhabi Emirate , Al Al-‘Ayn ( Gillett 1995 ). Fujairah Emirate , Fujairah . Ra’s al Khaymah Emirate , Khor Khuwair ( Král & Batelka 2017 ). Material examined . SAUDI ARABIA : Asir Province , Abha , Raydah Nature Reserve , 1772 m (1811.618′N 4223.420′E), 30.i.2015 , H. Al Dhafer , H. Fadl , M. Abdel-Dayem , A. El Torkey & A. El Gharbawy leg. 1 ♀ , 1 ♂ ( KSMA ) ; Magardah , Wadi Al Talalieh , 286 m (195.192′N, 4147.784′E), 1.vi.2012 , H. Al Dhafer & M. Abdel- Dayem , 1 ♀ , 2 ♂♂ ( KSMA ) ; ibidem , 1.vi.2012 , H. Al Dhafer , M. Abdel-Dayem , B. Kondratieff & A. Al Ansi leg. 1 ♀ ( KSMA ) ; Saloos Almanzar , Wadi Baqrah , 422 m (1847.511′N 4201.090′E), 4.xi.2013 , H. Al Dhafer , H. Fadl , A. El Gharbawy & I. Rasool leg. 4 ♀♀ , 2 ♂♂ ( KSMA ) ; ibidem , 4.xi.2013 , H. Al Dhafer , H. Fadl , M. Abdel-Dayem & A. El Torkey leg. 5 ♀♀ , 3 ♂♂ ( KSMA ) ; ibidem , 433 m (1847.565′N 4201.117′E), 31.v.2012 , M.S. Abdel-Dayem leg. 6 ♀♀ , 3 ♂♂ ( KSMA ) ; 10 km N Wadi Reem , 200 m (1755′28″N 4215′20″E), 30.iii.2017 , G. Magnani & D. Baiocchi leg. 1 ex. ( SZCM ) ; Wadi Reem , 136 m (1752.551′N 4216.664′E), 9.ii.2016 , H. Fadl leg. 2 ♂♂ ( KSMA ) ; ibidem , 325 m (1803′N 4214′E), 9.ii.2016 , J. Bezdĕk & D. Král leg. 4 exx. ( NMPC ) . Jizan Province , Abu Arish , Ali Abu Ashwari Farm , 73 m (1659.923′N 4250.059′E), 10.v.2018 , H. Al Dhafer et al. leg. 1 ♀ ( KSMA ) ; Al Kwamlah , 33 m (1657.978′N 4240.122′E), 11.v.2018 , SW, H. Al Dhafer et al. leg. 2 ♀♀ ( KSMA ) . Makkah Province , Wadi Gounonah , 348 m (1924.673′N 4136.388′E), 11.iii.2012 , M. Abdel-Dayem & A. El Torkey leg. 1 ♂ ( KSMA ) . OMAN : Ad Dhahirah Governorate , Near Mahdah (2422′N 5556′E), 25.ii.1992 , M. P. T. Gillett leg. 3 ex. ( MGCB ) ; ibidem , 17.iv.1992 , M. P. T. Gillett leg. 3 ex. ( MGCB ) ; ibidem , 14.v.1992 , M. P. T. Gillett leg. 2 ex. ( MGCB ) . Al Batinah Governorate , near Sohar (2418′17′′N 5644′46′′E), 13.x.95 , M. Balkenohl & M. Gallagher leg. 1 ex. ( MBPC ) . Al Buraimi Governorate , J. Huwayya (2418′N 5551′E), 10.iv.1992 , M. P. T. Gillett leg. 2 exx. ( MGCB ) . Ash Sharqiyah Governorate , Wahiba Sands (2139′N 5918′E), 6/ 7.vii.1995 , B. Skule & M. D. Gallagher leg. 3 exx. ( SOIC , EBCT ) . Dhofar Governorate , Ain Garziz , 140 m (1706′34′′N 5404′46′′E), 8.x.2001 , M. Dellacasa leg. 1 exx. ( MSNC ) ; ibidem , 2.x.2001 , M. Dellacasa leg. 3 exx. ( MSNC ) ; Ain Razat , 110 m (1707′75′′N 5414′24′′E), 21.ix.2001 , M. Dellacasa leg. 4 exx. ( MSNC ) ; ibidem , 22.ix.2001 , M. Dellacasa leg. 2 exx. ( MSNC ) ; ibidem , 27.ix.2001 , M. Dellacasa leg. 3 exx. ( MSNC ) ; ibidem , 29.ix.2001 , M. Dellacasa leg. 2 exx. ( MSNC ) ; ibidem , 2.x.2001 , M. Dellacasa leg. 7 exx. ( MSNC ) ; ibidem , 6.x.2001 M. Dellacasa leg. 2 exx. ( MSNC ) ; ibidem , 7.ix.2002 , M. Dellacasa leg. 1 exx. ( MSNC ) ; Al Mughsayl dint., 30 m (1653′00′′N 5346′50′′E), 2.x.2001 , M. Dellacasa leg. 35 exx. ( MSNC ) ; ibidem , 5.x.2001 M. Dellacasa leg. 13 exx. ( MSNC ) ; ibidem , 30 m (1653′01′′N 5346′47′′E), 23.ix.2001 , M. Dellacasa leg. 93 exx. ( MSNC ) ; ibidem , 27.ix.2001 , M. Dellacasa leg. 33 exx. ( MSNC ) ; ibidem , 6.ix.2002 , M. Dellacasa leg. 35 exx. ( MSNC ) ; ibidem , 9.ix.2002 , M. Dellacasa leg. 44 exx. ( MSNC ) ; ibidem (1653′02′′N 5347′70′′E), 11.ix.2000 , M. Dellacasa leg. 40 exx. ( MSNC ) ; ibidem (1653′83′′N 5346′31′′E), 6.ix.2002 , M. Dellacasa leg. 12 exx. ( MSNC ) ; ibidem , 12.ix.2002 , M. Dellacasa leg. 21 exx. ( MSNC ) ; ibidem (1653′84′′N 5346′35′′E), 2.ix.2002 , M. Dellacasa leg. 11 exx. ( MSNC ) ; ibidem (1653′74′′N 5348′97′′E), 28.viii.2000 , M. Dellacasa leg. 65 exx. ( MSNC ) ; ibidem , 5.ix.2000 , M. Dellacasa leg. 29 exx. ( MSNC ) ; ibidem , 8.ix.2000 , M. Dellacasa leg. 163 exx. ( MSNC ) ; Ain Sahanawt , env. (1707′40′′N 5409′44′′E), 24.ix.2001 , M. Dellacasa leg. 4 exx. ( MSNC ) ; ibidem (1708′60′′N 5410′87′′E), 1.x.2001 , M. Dellacasa leg. 6 exx. ( MSNC ) ; ibidem , 4.x.2001 , M. Dellacasa leg. 11 exx. ( MSNC ) ; ibidem , 7.x.2001 , M. Dellacasa leg. 5 exx. ( MSNC ) ; ibidem (1708′63′′N 5410′87′′E), 8.x.2001 , M. Dellacasa leg. 1 ex. ( MSNC ) ; Darbat , pool, 220 m (1705′70′′N 5426′80′′E), 22.ix.2001 . M. Dellacasa leg. 1 ex. ( MSNC ) ; ibidem , 1.ix.2002 , M. Dellacasa leg. 10 exx ( MSNC ) ; Kara Mts. , E of Mugurah , 396 m (1645,91′N 5328,00′E), 15.x.2009 , S. Ilniczky leg. 2 exx. ( SICB ) Kara Mts. , S of Mirbat , 95 m (1702,68′N 5432,345′E), 16.x.2009 , S. Ilniczky leg. 3 exx. ( SICB ) ; Kara Mts. , SW of Al Mughsayi , 418 m (1651,24′N 5343,149′E), 16.x.2009 , S. Ilniczky leg. 1 ex. ( SICB ) ; Khor Rori , 37 m (1702′27″N 5426′16″E), 24.viii.2000 , M. Dellacasa leg. 247 exx. ( MSNC ) ; ibidem , 4.ix.2000 , M. Dellacasa leg. 20 exx. ( MSNC ) ; ibidem , 12.ix.2000 , M. Dellacasa leg. 88 exx. ( MSNC , SZCM ) ; ibidem , 1.ix.2002 , M. Dellacasa leg. 6 exx. ( MSNC ) ; Rakhyut , env. (1645′00′′N 5324′67′′E), 12.ix.2002 , M. Dellacasa leg. 26 exx. ( MSNC ) ; Road Marbat-Sadh , km 30 E Marbat , 170 m (1704′67′′N 5453′39′′E), 6.x.2001 , M. Dellacasa leg. 3 exx. ( MSNC ) ; Road Marbat-Sadh , 134 m (1703′06′′N 5450′97′′E), 7.ix.2000 , M. Dellacasa leg. 34 exx. ( MSNC ) ; Road Salalah-Darbat , 23 m (1703′14′′N 5425′58′′E), 9.ix.2000 , M. Dellacasa leg. 13 exx. ( MSNC ) ; Road 41, between Raysut & Adwnab , 90 m (1657′72′′N 5352′73′′E), 23.ix.2001 , M. Dellacasa leg. 40 exx. ( MSNC ) ; Road 45, Ra- khyut env., 30 m (1645′07′′N 5324′59′′E), 30.ix.2001 , M. Dellacasa leg. 3 exx. ( MSNC ) ; Road 47, Adwnab , env., 113 m (1655′34′′N 5350′56′′E), 5.ix.2000 , M. Dellacasa leg. 14 exx. ( MSNC ) ; Road 47, Al Mughsayl , env., 30 m (1653′71′′N 5349′01′′E), 23.ix.2001 , M. Dellacasa leg. 11 exx. ( MSNC ) ; ibidem , 2.x.2001 , M. Dellacasa leg. 68 exx. ( MSNC ) ; ibidem , 5.x.2001 , M. Dellacasa leg. 5 exx. ( MSNC ) ; Road 47, ante Al Mughsayl , 60 m (1654′11′′N 5349′20′′E), 11.ix.2000 , M. Dellacasa leg. 18 exx ( MSNC , SZCM ) ; ibidem (1653′75′′N 5348′96′′E), 2.ix.2002 , M. Dellacasa leg. 92 exx. ( MSNC ) ; Road 49 near Ain Sahanawt (1707′40′′N 5409′40′′E), 24.ix.2001 , M. Dellacasa leg. 9 exx. ( MSNC ) ; Salalah , env., 58 m (1707′14′′N 5409′46′′E), I. Zappi leg. 5 exx. ( IZCC ) ; Salalah , Wadi Darbat , 41 m (1704′23”N 5425′56”E), I. Zappi leg. 6 exx. ( IZCC ) ; Taqah. env., 23 m (1703′14′′N 5425′58′′E), 26.viii.2000 , M. Dellacasa leg. 3 exx. ( MSNC ) ; ibidem , 4.ix.2000 , M. Dellacasa leg. 13 exx. ( MSNC ) ; Wadi Ashawq , 42 m (1653′88′′N 5346′31′′E), 5.ix.2000 , M. Dellacasa leg. 55 exx. ( MSNC ) ; ibidem , 11.ix.2000 , M. Dellacasa leg. 10 exx. ( MSNC ) ; ibidem , 40 m (1653′83′′N 5346′31′′E), 23.ix.2000 M. Dellacasa leg. 21 exx. MSNC ); Wadi Darbat (1704′50′′N 5425′88′′E), 24.viii.2000 , D. Gianasso leg. 1 ex. ( MSNC ) ; ibidem , 9.ix.2000 , D. Gianasso leg. 3 exx. ( MSNC ) ; ibidem , 7.ix.2002 , M. Dellacasa leg. 1 ex. ( MSNC ) ; ibidem , 9.ix.2000 , M. Dellacasa leg. 13 exx. ( MSNC ) . UNITED ARAB EMIRATES : Abu Dhabi Emirate , Ain Al Fayda (2406′N 5540′E), 7.v.1992 , M. P. T. Gillett leg. 5 exx. ( MGCB ) . FIGURES 8–13 . Onthophagini of Arabian Peninsula. 8, Onthophagus ( Furconthophagus ) aethiopicus d’Orbigny, 1902 : YE (Sana’a) Alharf; 9, Onthophagus ( Furconthophagus ) frontalis Raffray, 1877 : YE (Sana’a) Wadi Dhahr; 10, Onthophagus ( Furconthophagus ) rugulipennis Fairmaire, 1887 : YE (Sana’a) Wadi Dhahar; 11, Onthophagus ( Furconthophagus ) sellatus Klug, 1845 : SA (Riyadh) Deirab; 12, Onthophagus ( Furconthophagus ) variegatus ( Fabricius, 1798 ) : YE (Sana’a) Wadi Dhahr; 13, Onthophagus ( Indonthophagus ) nitidulus Klug, 1856 : OM (Dhofar) Salalah. Photos by Gabriele Fiumi. Distribution . Arabian Peninsula: Saudi Arabia ( Gestro 1889 ), United Arab Emirates ( Gillett 1995 ), Yemen (Scott in Paulian 1948 ). North Africa: Egypt ( Baraud 1985 ). Africa: Eritrea , Ethiopia , Somalia , Sudan ( Balthasar 1963 ). Asia: Afghanistan , Iran , Iraq , Pakistan ( Ziani & Bezděk 2016 ). Remarks . First record from Oman . Kabakov (2006) placed the species in the subgenus Indonthophagus . The subgeneric placement of Kabakov is tentatively followed here.