A study of the genus Protosticta Selys, 1855, with descriptions of four new species from Vietnam (Odonata: Platystictidae) Author Phan, Quoc Toan Author Kompier, Tom text Zootaxa 2016 4098 3 529 544 journal article 10.11646/zootaxa.4098.3.6 687e2f7e-8268-4707-9cd7-bf46f9753f3b 1175-5326 257723 A005DB6C-7389-4506-B2D5-392531F832EB 6. Protosticta satoi Asahina, 1997 —dark form ( Figure 5 A–K) Protosticta khaosoidaoensis satoi : Asahina (1997) , pp. 108–109, Figs. 3–4 . Type locality: Tam Dao National Park, Vinh Phuc Province, northern Vietnam .— Tsuda (2000) , p. 326 (catalogued Vietnam ). Protosticta khaosoidaoensis nec Asahina, 1997: Do & Dang (2007) , p. 65 (distribution map). Protosticta satoi : van Tol (2008) (new status), pp. 222–227, Figs. 11–16.— Do (2011) (serial photographs of emergence), pp. 5– 7, Figs. 1–9 .—von Ellenrieder et al. (2015) , p. 8, Fig. 12. Specimens examined . 5 males , 1 female , Pia Oac Nature Reserve, Cao Bang Province, 18.V.2015 , Quoc Toan Phan leg.; 1 male , Tam Dao National Park, Vinh Phuc Province, 18.VIII.2013 ; 4 males , Ba Be National Park, Bac Kan Province, 26.VII.2013 and 4 males , same location, 22.VI.2014 , Tom Kompier leg. Notes . Asahina (1997) described the subspecies P. khaosoidaoensis satoi (= P. satoi light form) on the basis of a single female from Tam Dao National Park (Vinh Phuc Province, northern Vietnam ). Van Tol (2008) subsequently elevated P. s a t o i as a distinct species from P. khaosoidaoensis based on specimens from Cuc Phuong National Park (Ninh Binh Province) and Tam Dao. As in P. beaumonti , there are no obvious structural differences between the dark and light froms of P. s at oi , but the forms are easily separated by their thoracic color pattern ( Figs. 5 A, 5H, 6A, 6C). Van Tol (2008) described the color pattern of the prothorax of the male P. s a t o i dark form from Tam Dao National Park as “anterior lobe black, median lobe with paired protuberance brownish black, medially darker”. This character is consistent with all dark form males of P. sat oi from several locations in northern Vietnam ( Figs. 5 B, 5I ), apart from one male from the same location (Tam Dao), which has the median lobe yellowish, anterior and posterior lobes darkened ( Fig. 5 K). Measurements . Male. Hindwing 19–22.5; abdomen including appendages 34–42. Female. Hindwing 22, abdomen including appendages 33–34.