Review of Pennella Oken, 1816 (Copepoda: Pennellidae) with a description of Pennella benzi sp. nov., a parasite of Escolar, Lepidocybium flavobrunneum (Pisces) in the northwest Atlantic Ocean Author Hogans, W. E. text Zootaxa 2017 4244 1 1 38 journal article 36256 10.11646/zootaxa.4244.1.1 05f1a37b-29f3-4bca-a735-178a3373bb8a 1175-5326 400400 781D71C8-4632-4D1B-8D82-F77CA1146029 Pennella makaira Hogans, 1988 ( Fig. 14 ) Synonyms. None Type host and locality. Makaira nigricans , Caribbean Sea. Morphology. Size: 27–29 mm . Papillae: full coverage, spherical, clavate and tubiform, simple branching; unorganized arrangement. Holdfasts: two only, short. First antenna with three segments, second with two segments. Plumes: simple, single or double. Remarks. A valid species. Found so far only on blue marlin ( M. nigricans ) in the south Atlantic ( Hogans 1988c ; Williams & Bunkley-Williams 1996 ). A small species with a unique overall shape and uniform papillae structure between specimens. The only small (< 50 mm ) Pennella known to infest large scombriform fishes.