The Afrotropical Robber Fly Genus Congomochtherus Oldroyd, 1979 (Diptera: Asilidae: Asilinae)
Author
Jason G. H. Londt
text
African Invertebrates
2014
2014-12-31
55
2
333
350
journal article
28882
10.5733/afin.055.0204
32cb17b8-b7f1-4cc9-9646-316f97621f29
1472896
B6764F23-CEA3-46CD-9CF5-56843CD40D99
Heligmonevra russnota
Martin, 1964
Stat. Rev
.
Figs 12-15
Heligmoneura russnota
Martin, 1964
: 304
(figs 73 mesonotum, 83 ♂ terminalia).
Heligmonevra russnota
: Oldroyd 1980: 338
(catalogue).
Congomochtherus russnota
:
Tomasovic 2006
: 148
(figs 6 epandrium, 7 gonocoxite & dististylus, 8 hypandrium, 9 aedeagus).
Martin (1964)
described this Madagascan species from the type locality of Ambalavao [
c
. 21°14'S 47°14'E
c
. 1280 m] and five other localities on the island. In the absence of a modern review of
Heligmonevra
,
Tomasovic (2006)
transferred
H. russnota
to
Congomochtherus
after studying material (9♂ 9♀) from Morarano Chrome, collected by A. Pauly and housed in GULB.As this was the first record of the genus from Madagascar, I borrowed two pairs of specimens from GULB in order to satisfy my curiosity regarding Tomasovic's action. While I can confirm the specific identification to be
H. russnota
, as the males possess the hypandrium uniquely tapering to a point (
Figs 12, 14
), I am not convinced that the species is correctly assigned to
Congomochtherus
and so here reassign it to
Heligmonevra
, where I believe it should reside, at least until a thorough revision of the genus has been undertaken. The Morarano Chrome specimens key out well to
Heligmonevra
using a key to the genera of Asilinae published by Londt (2002). There are currently 24 described Afrotropical species, 14 of which are Madagascan endemics (
Note
: Martin's (1964) study included 16 species, two of which have since been transferred to other genera).
Figs 12-15.
Heligmonevra russnota
Martin, 1964
: (12-14) ♂ terminalia (lateral, dorsal, ventral); (15) ♀ terminalia (lateral) (Scale bars = 1 mm).
In general appearance
Congomochtherus
species are far more robust and darkly pigmented than
Heligmonevra
species, which are rather slender and much paler in colour. While these genera are similar, the specimens of
H. russnota
do not agree with
Congomochtherus
species in a variety of features: (1)
In
H. russnota
the antennal style appears 2-segmented -the small basal segment-like element is not clearly defined (in
Congomochtherus
this element is reasonably distinct). (2)
In
H. russnota
all femora are slender and predominantly yellowish except for small dark-brownish areas (in
Congomochtherus
all femora are fairly robust and entirely to very extensively blackish). (3)
In
H. russnota
the thoracic pruinescence is strongly developed (in
Congomochtherus
species it is weakly developed). (4)
In
H. russnota
the face is only slightly ventrally protuberant (in
Congomochtherus
the face is obviously protuberant). (5)
In
H. russnota
the female terminalia (
Fig. 15
) are relatively weakly sclerotised, in particular the hypogynial valves, and cerci are finely setose (in
Congomochtherus
the female terminalia are strongly sclerotised and the cerci commonly have short spine-like setae).
Material examined: MADAGASCAR: 2♂ 2♀ Morarano [19°26'S 47°29'E
c
. 1550 m] - Chrome, i.1992, A Pauly (GULB).