Synopsis of the Grenadier Fishes (Gadiformes; Teleostei) of Taiwan
Author
Iwamoto, Tomio
Section of Ichthyology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA;
Author
Nakayama, Naohide
Laboratory of Marine Biology, Faculty of Science, Kochi University, 2 - 5 - 1 Akebono-cho, Kochi, 780 - 8073, Japan;
Author
Shao, Kwang-Tsao
Research Center for Biodiversity, Academia Sinica No. 128, Sec. 2, Academia Road, Nankang, Taipei 115, Taiwan;
Author
Table, Hsuan-Ching Ho
text
Proceedings of the California Academy of Sciences
2015
2015-04-15
62
3
31
126
journal article
299670
10.5281/zenodo.11512126
522b8de7-880d-40ca-803e-aa5a3a05c5f6
0068-547X
11512126
Bathygadus garretti
Gilbert and Hubbs, 1916
Bathygadus garretti
Gilbert and Hubbs, 1916:151–153, pl.8, fig. 1 (
holotype
USNM 76863
,
513
TL,
Suruga Gulf
,
Japan
,
Albatross
sta. 5059, 197–297 fm [
360–543 m
])
.
Bathygadus
(
Melanobranchus
)
garretti
: Gilbert and Hubbs, 1920:380 (in key).
Bathygadus garretti
:
Okamura, 1970:34–36
, pl. 12, text-fig. 17 (descr. based on
paratype
,
USNM
135351);
Okamura, 1984:197
, 357, fig. 139 (descr. in Japanese and English;
7 spec.
,
Okinawa
Trough,
360–650 m
).—
Chiou et al., 2004b:42
, fig. 6 (
1 spec.
, Da-xi,
NET
).—
Shao et al., 2008
: table 2 (
6 spec.
,
NET
,
SWT
,
SCS
).
Bathygadus nipponicus
: Howes and Crimmen, 1990:191 (in part; synonymized
B. garretti
with
B. nipponicus
).
Chiou et al., 2004b:42
, fig. 7 (
1 spec.
,
NET
).
MATERIAL
EXAMINED
(
20 spec.
).—
NET
:
ASIZP 61226
(1, 86.5 HL, 446 TL)
; Da-xi;
ASIZP 61227
(2, 70–77 HL, 362–380 TL)
; Da-xi;
ASIZP 61228
(1, 96.8 HL, 458 TL)
; Da-xi,
ASIZP 64274
(1, 91.3 HL, 365 TL)
; CP
235, 764 m
;
ASIZP 70241
(1, 281 TL)
; Da-xi;
ASIZP 70244
(1,
153 TL);
Da-xi
;
ASIZP 70246
(3, 127–175 TL);
Da-xi
;
ASIZP 70249
(3, 360–456 TL);
Da-xi
;
ASIZP 70253
(1, 444 TL);
Da-xi.
SWT
:
ASIZP 65512
(1, 58.1 HL, 320 TL); CP 130, 709–
728 m
;
ASIZP 65593
(1, 70 HL, 338 TL); CD 140, 280–
452 m
;
ASIZP 65599
(1, 65.6 HL, 291+ TL); CD 137, 316–
477 m
.
No
data:
ASIZP 65580
(1, 71.6 HL, 335+ TL).
Other
material
:
Japan
:
BSKU 98225
(1, 62.5 HL, 305+ TL) and
BSKU 98224
(1, 58.9 HL, 320 TL);
Suruga Bay
,
520–545 m
;
23 Nov. 1978
.
DISTINGUISHING
FEATURES
.— A species of
Bathygadus
with gular membrane scaled; a rudimentary chin barbel usually present (sometimes absent); dorsal profile behind head relatively low, nape not humpbacked; 1D II8–10; P i15–i18;
V
10 (rarely 11); GR-I (5–6) + (17–20); pyl. caeca 34–46 (or more); interorbital width 27–32% HL; orbit diameter 21–26%; upper jaw length 52–58%; none of fins with greatly elongated rays.
DISTRIBUTION
.—
Southern
Japan
s. to
Taiwan
(
NET
,
SWT
) in
360–650 m
.
REMARKS
.— Our identification of specimens of this species was based on a combination of characters, including the presence of a rudimentary chin barbel. In a few specimens, the barbel was not present, but other characters that we deemed important appeared to confirm our identification. Howes and Crimmen (1990:1910) synonymized
B. garretti
with
B. nipponicus
, stating that the only difference between the two was the presence of a rudimentary chin barbel in the former, a suggestion made earlier by
Okamura (1984:358)
. However, our
ASIZ
specimens suggest that
B. garretti
almost always has
V
10 (11 on one side in a
paratype
) vs. mostly
9 in
B. nipponicus
(the
holotype
has 10,
fide
Okamura 1970
and Howes and Crimmen 1990, not 9 as stated in the original description), a slightly larger orbit (1–26% HL vs. 17–22%), and more pyloric caeca (34–46 [
5 spec.
] vs. 12–19 [
5 spec.
]). The count of pyloric caeca in the single
paratype
of
B. garretti
was given as 50 by Gilbert and Hubbs (1916:152), 49 by
Okamura (1970:34)
, and 54 by Howes and Crimmen (1990:191).
Specimen
ASIZP
61227, listed in
Chiou et al. (2004b)
as the first
Taiwan
record of
B. nipponicus
, was determined by us to be
B. garretti
. One of
two specimens
of
B. antrodes
(
ASIZP
61226) recorded in the same paper is now re-identified as this species.