A survey of Palaearctic Dictynidae (Araneae). 1. Taxonomic notes on Dictynomorpha Spassky, 1939, Brigittea Lehtinen, 1967 and Lathys Simon, 1884
Author
Marusik, Yuri M.
Author
Esyunin, Sergei L.
Author
Tuneva, Tatyana K.
text
Zootaxa
2015
3925
1
129
144
journal article
10.11646/zootaxa.3925.1.9
98786b6c-26a1-4204-9967-e2190ecce616
1175-5326
243577
52C2B852-7814-4313-B18A-717241C81E4F
Dictynomorpha
Spassky, 1939
Dictynomorpha
Spassky 1939
: 138
.
Dictynomorpha
:
Lehtinen 1967
: 230
, 360.
Type
species
:
D. strandi
Spassky, 1939
from
Kyrgyzstan
, by monotypy.
Comments
. The genus was originally described for one species from
Kyrgyzstan
,
Dictynomorpha strandi
Spassky, 1939
.
Lehtinen (1967)
transferred three additional species to
Dictynomorpha
, without studying the
type
material of
D. strandi
and
types
or comparative specimens of other species placed in the genus. Currently, four species are assigned to this genus:
D. bedeshai
(Tikader, 1966)
,
D. marakata
(Sherriffs, 1927)
(both from
India
),
D. smaragdula
(
Simon, 1905
)
(
Sri Lanka
) and
D. strandi
(Central Asia). The diagnosis of the genus is based mainly on the male palp which has a strongly modified patella and cymbium. Hence, it is unclear why Lehtinen transferred
D. bedeshai
, a species known only from females into
Dictynomorpha
. Additionally, the male palp of
D. smaragdula
differs significantly from the palp of the generotype by having a small ventral process of the patella (
D. strandi
has a large dorsal process) and lacks a protruding extension of the cymbium, which is strongly protruding in
D. strandi
. It appears likely that all species from
India
and
Sri Lanka
are misplaced in the genus.
Relationships
. Judging from the habitus and male palp, the genus is related to
Ajmonia
Caporiacco, 1934
, a taxon known from the South Palaearctic.
Dictynomorpha
is also somewhat similar to the Oriental
Anaxibia pictithorax
(Kulczyński, 1908)
and Neotropical
Thallumetus
Simon, 1893
because all three of these taxa have modified patella, tibia and cymbia. The male of the
type
species of
Anaxibia
Thorell, 1898
is unknown. These four genera have not been revised and can not be compared in any detail with
Dictynomorpha
. Copulatory organs are relatively well studied only in Oriental
Ajmonia
(
Marusik & Esyunin 2010
)
.
Diagnosis
.
Dictynomorpha
can be distinguished by the modified patella, tibia and cymbium of the male palp.
Dictynomorpha
differs from the related
Ajmonia
by having the male palpal femur modified with a small knob (unmodified in
Ajmonia
), and a strongly developed processes on the patella and cymbium (
vs
. one process on patella and usually one considerably smaller process/extension of cymbium in
Ajmonia
).
Composition
. Two species can be considered in the genus:
D. strandi
and
D. daemonis
sp. n.
Here, we transfer
D. smaragdula
(cf.
Figs 14–15
) to
Ajmonia
[
Ajmonia smaragdula
(
Simon, 1905
)
comb. n
.
], based on the modified cymbium and tibia. We also transfer two other species,
Dictynomorpha bedeshai
(Tikader, 1966)
and
D. marakata
(Sherriffs, 1927)
, to
Ajmonia
.
However, this is a tentative placement because the former species is known only from a female, and the latter has no proper figures but they do not belong in the genus
Dicytnomorpha
[
Ajmonia bedeshai
(Tikader, 1966)
comb. n
.
and
Ajmonia marakata
(Sherriffs, 1927)
comb. n
.
].