The genus Diploneis Ehrenberg ex Cleve (Bacillariophyta) from Lake Hövsgöl, Mongolia
Author
Jovanovska, Elena
Department of Animal Ecology and Systematics, Justus-Liebig University, Heinrich-Buff-Ring 26 - 32, 35392 Giessen, Germany
jovanovska.eci@gmail.com
Author
Levkov, Zlatko
Institute of Biology, Faculty of Natural Sciences, Arhimedova 3, 1000 Skopje, R. of Macedonia
Author
Edlund, Mark B.
St. Croix Watershed Research Station, Science Museum of Minnesota, Marine on St. Croix, Minnesota 55047, USA
text
Phytotaxa
2015
2015-06-26
217
3
201
248
http://dx.doi.org/10.11646/phytotaxa.217.3.1
journal article
10.11646/phytotaxa.217.3.1
1179-3163
13634131
Diploneis exigua
sp. nov
.
(
Figs 152–165
)
Valves are elliptical with convex margins and round ends (
Figs 152–160
). The valve length is 14.5–17.0 μm, and the valve breadth is 9.0–10.5 μm. The axial area is narrow, linear, and expands into a small rectangular central area. From inside, a thin silica plate covers the whole length of the longitudinal canal (
Fig. 165
). From outside, the central area is small and rectangular, ca. 1.5 μm wide. Externally, the longitudinal canal appears linear with one to two rows of small areolae in the middle of the valve continuing into one or two areolae covered with a large cribrum towards the valve apices (
Figs 160, 161
). The areolae of the longitudinal canal are covered with cribra morphologically identical to those on the striae, from which they are separated with a narrow hyaline area (
Fig. 161
). In the middle of the valve the areolae of the canal are covered with small and simple cribra that become larger and irregularly polygonal towards the valve apices (
Fig. 160
). Internally the longitudinal canal is closed with a thick linear silica plate forming a “depression” where the raphe is placed (
Fig. 165
). Externally, the raphe is straight and simple with expanded drop-like proximal ends that are bent to the same side of the valve and positioned within an expanded depression (
Fig. 160
). Toward the central area the raphe branches drop into a “depression” below the rest of the non-porous silica, and are surrounded with a thick silica ridge (
Fig. 160
). The depression is deepest and widest at the central area and gradually narrows toward the raphe distal ends. Distally, the raphe branches are bent into a short drop-like terminal fissures deflected to the same side of the valve (
Figs 160, 161
). Internally the raphe is straight with simple proximal ends, inserted in a slightly elevated sternum inside the “depression” formed by the longitudinal canal (
Fig. 165
). Distally, the raphe endings are slightly bent to the same side of the valve into very short terminal fissures (
Fig. 163
). Striae are parallel in the middle of the valve, becoming radiate at the distal ends of the valve,
15–16 in
10 μm, and composed of round areolae,
16–20 in
10 μm. Striae are uniseriate becoming biseriate and/or complex in structure towards the valve margins. From outside, the areolae are covered with cribrate occlusions, which increase in size towards the valve margins, becoming largest at the mantle (
Figs 160–162
). Internally the alveoli open via single continuous and elongate openings covered with a fine silica layer (
Figs 162–165
).
Type:—
MONGOLIA
,
Lake
Hövsgöl
(
Hövsgöl
National Park
),
Station
anchored off
Hanh Gol
mouth.
Coordinates
:
51°27.268’ N
;
100°43.11’E
,
Chara
from
4 m
depth
(accession number: M272
A, M.B. Edlund Collection
,
Science Museum of Minnesota
, collected by
Mark B. Edlund
and
Nergui Soninkhishig
,
19 July 1998
) (
Slide
M272A,
ANSP
GC-36355
, GCM-24057),
holotype
, designated here
; example specimens on
Figs 153, 156
;
(
Slide
919072,
CAS
,
isotype
designated here
)
.
Etymology:—
The species name refers to the taxon’s small size.
Observations:
—With respect to the valve size,
Diploneis exigua
is very similar to
D
.
soninkhishigae
, but differs in: valve shape (elliptical
vs
. elliptical-lanceolate); central area (rectangular
vs
. elongate); striae density (
15–16 in
10 μm
vs.
11–15 in
10 μm); and areola density (
16–20 in
10 μm
vs
.
10–12 in
10 μm).
Diploneis exigua
has a similar central area as
D. stoermeri
morphotype 1 (compare
Figs 152–159
with
Figs 80–89
). However, the size range (length: 14.5–17.0 μm in
D
.
exigua
vs
. 19.5–44.5 μm
D
.
stoermeri
; breadth: 9.0–10.5 μm in
D
.
exigua
vs
. 12.5–22.0 μm in
D
.
stoermeri
) separates these two species.
Diploneis paraparma
differs from
D
.
exigua
by the greater size range (length: 19.5–42.0 μm
vs
. 14.5–17.0 μm; breadth: 12.5–16.5 μm
vs
. 9.0–10.5 μm) and the striae structure (uniseriate becoming biseriate towards the valve margins
vs
. uniseriate).
Ecology and Distribution:
—M248A; M272A; M273A; M274A; M291A: found in depths from
4 to 40 m
in the central and northern areas of Lake
Hövsgöl
in sediment and epiphytic on
Chara
.