Order Chiroptera - Family Hipposideridae Author Wilson, Don E. Author Reeder, DeeAnn text 2005 The Johns Hopkins University Press Baltimore Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1 365 379 book chapter 0-8018-8221-4 10.5281/zenodo.7316519 Hipposideros ater subsp. saevus K. Andersen 1918 Synonyms: Hipposideros ater subsp. toala Shamel 1940 . Discussion: bicolor species group.