Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru) Author DeVries, Thomas J. text Journal of Natural History 2019 2018-12-17 53 25 1533 1584 journal article 10.1080/00222933.2018.1524032 034ebcd9-efa7-416a-822c-b6aee0e57a1c 1464-5262 3670229 Genus Carolia Cantraine, 1838 Remarks The genus Carolia Cantraine, 1838 , is thought to have appeared for the first time in lower Eocene strata of Egypt, India, and Pakistan and to have spread across the Tethyan region during the middle Eocene ( El-Shazly et al. 2016 ). Specimens from Maastrichtian beds in Egypt identified as examples of Carolia ( Quaas 1902 ) were subsequently assigned to a new genus, Tarturia Strougo, 1983 ( Strougo 1983 ), thought to be ancestral to Carolia (El Shazly et al. 2016) . Species of Carolia have been described from lower and lower middle Eocene beds of India ( Cox 1938 ) and Oligocene beds in Argentina ( Ihering 1907 ).