Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae) Author Pall-Gergely, Barna https://orcid.org/0000-0002-6167-7221 Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary pallgergely2@gmail.com Author Hunyadi, Andras Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary Author Vermeulen, Jaap J. JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands Author Grego, Jozef Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia Author Sutcharit, Chirasak Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand Author Reischuetz, Alexander Puechhaimgasse 52, A- 3580 Horn, Austria Author Dumrongrojwattana, Pongrat Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand Author Botta-Dukat, Zoltan Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary Author Oerstan, Aydin 12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA Author Fekete, Judit University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary & Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary Author Jochum, Adrienne https://orcid.org/0000-0002-6624-6412 Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland & Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland & Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany text ZooKeys 2023 2023-02-13 1147 1 177 http://dx.doi.org/10.3897/zookeys.1147.93824 journal article http://dx.doi.org/10.3897/zookeys.1147.93824 1313-2970-1147-1 9BB9881B0076473D8E53155D37CA1F50 FF2B6B317B505F9EA0E1000BDCD16CE7 Angustopila papaver Pall-Gergely & Hunyadi sp. nov. Fig. 73 Type material. Holotype : Vietnam • 1 empty shell (H: 0.66 mm, D: 0.66 mm); Thanh Hoa Province, Như Thanh District, Hải Van , Hang Lo Cao Khang Chiến, vicinity of the cave (locality code: 2020/41); 19°37.08'N , 105°34.63'E ; 20 m a.s.l.; 14 Feb. 2020; A. Hunyadi leg.; HNHM 105297. Paratypes : Vietnam • 1 figured shell + 39 shells + 3 live-collected specimens covered in mud; same data as for holotype; coll. HA. Diagnosis. A small Angustopila species with a conical to conical-globular shell, a shouldered body whorl, a strong parietal tooth reaching the peristome edge, and an upper palatal tooth of ca. 1/2 size. Description. Shell small for the genus, shell slightly higher than wide or rarely as high as wide; off-white, translucent when fresh, conical to conical-globular; body whorl widest in apertural view; protoconch consists of 1-1.25 whorls with spiral striation preceding the first teleoconch whorl; teleoconch with very weak, regularly spaced radial ribs crossed by stronger spiral striae (ca. 12-15 on body whorl from apertural view); whorls ca. 3.5, slightly shouldered, "pushed" from baso-lateral direction from apertural view; aperture very slightly oblique to shell axis from lateral view; umbilicus moderately wide; aperture pear-shaped, sinulus narrow and strongly isolated due to the near joining of the strong parietal and upper palatal teeth; peristome expanded, not reflected; parietal callus slightly protruding, slightly detached from penultimate whorl (aperture not adnate); parietal tooth elevated, strong, short, reaches parietal callus, slightly curved in direction of palatal tooth; upper palatal tooth ca. 1/2 as high as parietal tooth, situated near peristome edge, just opposite of parietal tooth. Measurements (in mm). H = 0.66-0.75, D = 0.64-0.75, H/D*100 = 100-112.5 ( n = 7), RUD = 26.6-29.2 ( n = 3). Differential diagnosis. Similar only to A. coprologos and A. psammion in size. This new species can be distinguished from all congeners based on the combination of the very small, conical to conical-globular shell and the two apertural denticles. Etymology. The tiny shells of this species with approximately equal shell height and width resemble poppy ( Papaver ) seeds. The specific epithet is to be used as a noun in apposition. Figure 73. Angustopila papaver Pall-Gergely & Hunyadi, sp. nov. A, C-I holotype (HNHM 105297) B paratype. Apertural ( A, B ), lateral ( C ), apical ( D ) and ventral ( E ) sides of the shell; sculpture on the protoconch ( F ); aperture ( G ); ventral ( H ) and frontal ( I ) surface of the body whorl. Distribution. Angustopila papaver sp. nov. is known from the type locality only in Thanh Hoa Province of northern Vietnam (Fig. 48 ).