Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera Author Yoshitomi, Hiroyuki text European Journal of Taxonomy 2019 2019-09-04 583 1 45 journal article 24720 10.5852/ejt.2019.583 104a1ea7-b7d6-45cd-9402-bc84ab16b143 3564532 4F834ADD-57E7-4CE3-B16A-127F1BE4AE14 Mexico masamii ( Satô, 1994 ) comb. nov. Figs 11F , 18–19 Babalimnichus masamii Satô, 1994: 175 . Babalimnichus masamii – Yoshitomi & Satô 2001: 472 [larval description]. — Yoshitomi 2014a: 29 . — Ono et al. 2012: 47 . — Kamezawa 2017: 54 . Material examined JAPAN • 7 ex. ; “ Japan : Honshû Nanaura-kaigan Tachibana , Sado-shi Niigata , 5. vi. 2016 37.986591 , 138.234143 Kamezawa H. leg.”; EUMJ • 1 ♂ ; “[IZ9] Ishijirogawa , Shikinejima , Izu Isls. , Japan , 34°19.409″ N, 139°13.308″ E, ca 16 m , 7. II. 2013 , H. Yoshitomi leg.”; EUMJ • 1 ♂ , 1 ♀ , 14 ex. ; “ Kariya-kaigan Tokoname-shi Aichi Pref. 17. VIII. 1995 , 17. VIII. 1995 H. Yoshitomi leg.”; 1 ♂ and 1 ♀ on slides HY 989, 992–994; EUMJ • 1 ex. ; “ Zoubigasaki , Murozumi Hikari-shi Yamaguchi Pref. , Jpn 28. VII. 2005 Y. Fujitani leg.”; EUMJ • 5 ex. ; “ Bashayamamura Amami-oshima 3-V-1999 M. Sato leg.”; EUMJ • 28 ex. ; “ Kawata Okinawa 5-V-1999 M. Sato leg.”; EUMJ • 9 ex. ; “ Kim-kaigan Okinawa 28- III-1997 M. Satô leg.”; EUMJ • 1 ex. ; “ Onnason Okinawa-jima 26-28. IV. 2004 M. Sato leg.”; EUMJ • 1 ex. ; “ Kume-jima Ryukyu 30-IV-2004 M. Sato leg.”; EUMJ • 2 ex. ; “ Tomori Ryukyus Miyako-jima 7-V-1998 M. Satô leg.”; EUMJ • 13 ex. ; “ Uganzaki Ishigaki 22-IX-2003 M. Sato leg.”; EUMJ • 7 ex. , in ethanol; “ Sakaura , Izumo , Shimane 31. VII. 2006 M. Hayashi ”; EUMJ • 2 pupae , with larval skins, in ethanol; “ Nahama-kaigan , Nagai , Yokosuka-shi Kanagawa Pref. , Japan 6. July 2007 M. Asano leg.”; EUMJ . Fig. 18. Mexico masamii ( Satô, 1994 ) , male (A–F) and female (G–J) genitalia (EUMJ). A, G . Sternite VII. B . Sternite VIII. C . Sternite IX. D . Aedeagus, ventral view. E . Aedeagus in dorsal view. F . Aedeagus in right lateral view. H . Urosternite. I . Ovipositor. Redescription Male For a detailed description of the external features, see the original description ( Satô 1994 ). PW/PL 1.67– 2.22 (1.99); EL/EW 1.27–1.48(1.36);EL/ PL 2.64–3.29 (2.96); EW/PW 1.00–1.18 (1.10);TL/EW1.71– 1.95 (1.82). Sternite VII ( Fig. 18A ) bearing about 30 long setae, lacking extra setae; postero-lateral corners short and pointed; median part of caudal margin gently projecting triangularly. Sternite VIII ( Fig. 18B ) small, slightly sclerotized, Y-shaped. Sternite IX ( Fig. 18C ) well sclerotized, curved in basal part, shallowly concave at apex; apical plates rather stout, pointed at apices. Aedeagus ( Fig. 18 D–F) relatively short, asymmetrical; basal piece oval; lateral lobes short, with 3–4 apical setae; median lobe wide, sparsely punctuate in apical part, expanded in apical quarter, gently pointed at apex; ventral plates wide, separated from median lobe, left one long and curved interiorly in apical part, right one short and straight; ML/BL 2.24; ML/LL 2.43. Female Sexual dimorphism slight. Sternite VII ( Fig. 18G ) similar to that of male. Urosternite ( Fig. 18H ) well sclerotized, T-shaped, with long and slender apodeme. Ovipositor ( Fig. 18I ) well sclerotized; coxite sparsely punctuate, rather pointed at apices; apex of baculus bifid; approximate ratio of coxite and baculus (n = 1) 1.0:5.9. Pupae Body ( Fig. 19 ) about 1.5 mm , cream-colored, bearing long setae on head, pronotum, elytra and abdomen. Fig. 19. Pupa of Mexico masamii ( Satô, 1994 ) . A . Ventral aspect. B . Dorsal aspect. Scale bar: 1.0 mm. Measurements Unsexed (n = 20): TL 1.82–2.32 (2.04) mm; PW 0.90–1.10 (1.02) mm; PL 0.45–0.60 (0.52) mm; EL 1.32–1.72 (1.53) mm; EW 1.00–1.30 (1.12) mm. Distribution Japan (Honshu, Sadogashima, Izu Islands (Shikine-jima, Kozu-shima), Shikoku, Kyushu, the Ryukyus (Yakushima, Amami-Ôshima, Tokuno-shima, Okinoerabu-jima, Yoron-tô, Okinawa-jima, Irabu-jima, Kume-jima, Miyako-jima, Ishigaki-jima)). Remarks I re-examined the specimens from Yonaguni-jima ( Yoshitomi & Arai 2004 ), and it is clear that they do not represent this species but the preceding one. This is the first record of this species from Kume-jima. This species has the northernmost distribution (Sado Island) in the subfamily Thaumastodinae ( Kamezawa 2017 ) . Biological notes Overwintering takes place in the larval stage in Chiba Prefecture ( Ono et al. 2012 ) or the adult stage on the Izu Islands ( Yoshitomi 2014a ).