Taxonomic and Zoogeographic Study of the Japanese Phygadeuontinae (Hymenoptera, Ichneumonidae), with Descriptions of 17 New Species Author Watanabe, Kyohei Kanagawa Prefectural Museum of Natural History, 499 Iryuda, Odawara, Kanagawa 250 – 0031, Japan watanabe-k@nh‛kanagawa-museum‛jp text Bulletin of the Kanagawa Prefectural Museum (Natural Science) 2021 2021-03-30 50 55 136 https://doi.org/10.32225/bkpmnh.2021.50_55 journal article 300009 10.32225/bkpmnh.2021.50_55 70224806-ca38-47ca-bd8a-63da3ac532f9 2189-6720 12774639 DD695C62-DC2F-4D32-8D33-52648E28661D Lochetica japonica sp. nov. (New SJN: Samehada-chibi-togari-himebachi) ( Figs 47A–H , 62J , 65Q , 66Q ) Etymology . The specific name is from Japan. Type series . Holotype : KPM-NK 81989 , F , JAPAN , Tochigi Pref. , Nasushiobara City , Shiobara , Utou-sawa , 22–28. V. 2008 , T. Matsumura leg. ( MsT ) . Paratype : JAPAN , KPM-NK 81988 , F , Hokkaido , Sapporo City , Mt. Soranuma-dake , 14. VI. 2007, 4. VII. 2007 , A. Ueda leg. ( MsT ) . Description . Female (n=2). Body length 5.2–5.5 (HT: 5.2) mm. Body polished, covered with dense silver setae. Head. Clypeus 1.8–2.0 (HT: 1.8) times as wide as maximum length, its anterior margin rounded and obtuse, with dense setae along anterior margin ( Fig. 62J ). Face densely punctate. The border between clypeus and face indistinct ( Fig. 47B ). MSL 0.7–0.8 (HT: 0.7) times as long as BWM. Frons, gena, and vertex densely punctate, inter space of punctures coriaceous. Gena strongly convex in dorsal view ( Fig. 47C ). OOL slightly shorter than POL. OOL and POL longer than OD. Occipital carina complete, its lower end joined with hypostomal carina distant from mandibular base. Mandible covered with dense setae and its ventral margin indistinct. Upper tooth of mandible as long as lower tooth. Base of mandible slightly convex. Antenna with 20–21 (HT: 20) flagellomeres. FL III 2.2 times as long as maximum depth in lateral view. Mesosoma. Lateral part of pronotum densely punctate, the punctures on ventral part more or less longitudinally lined. Epomia long below pronotal sulcus, its lower part parallel to front edge of pronotum. Mesoscutum matt, minutely punctate. Notaulus short and weak. Scutellum densely punctate, without a lateral longitudinal carina. Mesopleuron densely punctate except for small smooth areas below subtegular ridge and in posterior part of speculum, the punctures more or less longitudinally lined. Epicnemial carina strongly developed dorsally, reaching subtegular ridge. Posterior transverse carina of mesosternum largely absent in front of mid coxae. Metapleuron densely punctate, with a weak, partly indistinct juxtacoxal carina. Propodeal carinae complete ( Figs 47G , 65Q ). Area basalis indistinct anteriorly ( Figs 47G , 65Q ). Area superomedia present, pentagonal ( Figs 47G , 65Q ). Anterior part of propodeum densely punctate ( Fig. 47G ). Median part of propodeum irregularly rugulose ( Fig. 47G ). Posterior part of propodeum transversely rugulose, the interspace of rugae smooth. Length of fore wing 4.0–4.1 (HT: 4.0) mm. Areolet present, pentagonal ( Fig. 47F ). Vein 2m-cu of fore wing with a single bulla. Nervellus inclivous, intercepted behind the middle ( Fig. 47F ). Hind femur 3.6–3.7 (HT: 3.6) times as long as maximum depth in lateral view. Hind TS I: II: III: IV: V = 2.0: 0.9: 0.6: 0.3: 0.6. Tarsal claws simple. Metasoma. T I 1.9–2.0 (HT: 2.0) times as long as maximum width, densely punctate ( Fig. 47H ) except for smooth base. Median dorsal carina of T I absent. Dorsolateral carina of T I complete. T II almost as long as wide. T II to T V densely punctate ( Fig. 47H ). Ovipositor sheath 2.3–2.38 (HT: 2.38) times as long as hind tibia. Ovipositor straight, with a weak but distinctly angulated nodus and ventral teeth ( Fig. 66Q ). Fig. 47. Lochetica japonica sp. nov. , KPM-NK 81989, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, dorsal view; D, head, mesosoma, and metasoma, dorsal view; E, head and mesosoma, lateral view; F, wings; G, propodeum, dorsal view; H, metasoma, dorsal view. Coloration ( Figs 47A–H ). Body (excluding wings and legs) black to blackish brown. Ventral surfaces of scape and pedicel, base of FL I, and apical part of mandible tinged with reddish brown. Palpi and tegula dark yellowish brown. Posterior margins of T I to T IV narrowly tinged with reddish brown. Membranous part of metasomal sternites dark brown. Ovipositor yellowish brown. Wings hyaline. Veins yellowish brown to dark brown. Pterostigma dark brown. Legs blackish brown to black. Fore and mid legs except for coxae at least partly paler than other parts of legs. Hind tibial spurs yellowish brown. Male. Unknown. Distribution . Japan ( Hokkaido and Honshu). Bionomics . Unknown. Remarks . This species resembles L. westoni in the long epomia, the strongly bordered area postero-externa, and the distinct nodus with angulation, but can be distinguished by the above key.