The first fossil leptofoenine wasp (Hymenoptera, Pteromalidae): A new species of Leptofoenus in Miocene amber from the Dominican Republic Author Engel, Michael University of Kansas, Natural History Museum, Lawrence, United States of America text ZooKeys 2009 2009-07-03 13 13 57 66 journal article 10.3897/zookeys.13.159 3694a5b6-d12b-4bdf-ad8c-aea2000cec4e 1313–2970 576466 94FB118C-4FD6-49AB-ADA6-80313E2B6212 Leptofoenus pittfieldae Engel , sp. n. urn:lsid:zoobank.org:act: EA9AF9B8-EBBA-4F75-A447-99D0B2F3A3A5 Figs 3-5 Holotype . Male, KU-NHM-Ent DR-019; Early Miocene amber, Dominican Republic , La Toca group of mines northeast of Santiago , excavated in late Autumn 2008, Keith Luzzi coll.; deposited in the Insect Fossil Collection, Division of Entomology , University of Kansas Natural History Museum. Diagnosis . The new species can be readily recognized from its modern and mainland congeners by the combination of a thinly sclerotized band posterior to M+Cu in the forewing ( Fig. 5 ), the presence of a sclerotized spot in the forewing at the junction of M+Cu and the basal vein ( Fig. 5 ), M+Cu being comprised of a series of 14 setae, the basal cell containing 22 setae, the anterior margin of the costal cell sclerotized with a dense fringe of short fuscous setae, the hind wing with only three hamuli, the apicalmost antennal article apparently composite and resulting in an overall 11-segmented antenna, and the absence of a rasp-like structure on the inner surface of the metatibia. Description . Male: Total length 8.8 mm ; forewing length 4.7 mm ; head 1.0 mm in length; mesosoma 3.3 mm in length, petiole 1.2 mm in length; gaster 3.3 mm in length. Integument dark brown throughout ( Fig. 3 ). Head with scrobal cavity carinate laterally; maxillary palpus 4-segmented, apical palpomere elongate, with numerous elongate black setae dorsally; clypeus bare except for two erect setae laterally near oral fossa (visible in left, oblique view); antenna apparently 11-segmented, apical flagellar article elongate and apparently with lines indicating possible fusion of two very minute apical articles (resulting in an otherwise 13-segmented antenna); parascrobal crests and vertex strongly tranversely striate, forming rasp-like structure dorsally. Mesosoma with pronotum and mesoscutum strongly, transversely strigate ( Fig. 4 ), strigae increasingly bearing spicules posteriorly on mesoscutum; pronotum elongate, anterior margin rounded, posterolateral surface with striolate area ( Fig. 4 ), striolate area ovoid (distinctly longer than wide; more rounded in L . rufus LaSalle and Stage ), striations in striolate area finer and more evenly spaced anteriorly; mesoscutellum strongly transversely strigate, with numerous spicules; mesopleuron coarsely and irregularly punctate except in impressed region largely glabrous; propodeum laterally densely and contiguously punctate, with sparsely scattered setae. Procoxa with strong carina along dorsolateral margin; metacoxa sculptured as on lateral surface of propodeum; metafemur longitudinally striate ventrally except reticulate basally; metatibia without longitudinal rasp-like structure on inner surface; metabasitarsus twice as long as second tarsomere (tarsi pentamerous). Forewing with anterior margin of costal cell sclerotized ( Fig. 5 ), with dense fringe of short fuscous setae; basal cell with 22 setae; sclerotized spot at junction of M+Cu and basal vein ( Fig. 5 ); M+Cu composed of series of 14 setae and posteriorly by thinly sclerotized band leading to sclerotized spot ( Fig. 5 ); hind wing with three hamuli. Metasoma with petiole approximately 7 times longer than wide (direct dorsal view not possible so this is estimated); petiole weakly transversely strigate dorsally; gaster imbricate. Figure 3. Leptofoenus pittfieldae Engel sp. n. (KU DR-019), photomicrograph of male holotype (length of specimen 8.8 mm). Figures 4-5. Photomicrographs of holotype male of Leptofoenus pittfieldae Engel sp. n. (KU DR-019). 4 Detail of lateral surface of pronotum showing posterior striolate region. 5 Basal third of forewing showing sclerotized spot and distribution of setae along M+Cu and within basal cell. Etymology . The specific epithet is a matronym honoring Ms. Morgan Pittfield, enthusiast of all things adventurous when it comes to amber and niece of Keith Luzzi, who generously donated the holotype and permitted its study.