First record of the parasitoid family Ismaridae (Hymenoptera, Diaprioidea) from Eocene Baltic and Rovno ambers with the description of a new genus and two new species Author Chemyreva, Vasilisa G. Zoological Institute, Russian Academy of Sciences, 1 Universitetskaya Emb., St Petersburg 199034, Russia Author Perkovsky, Evgeny E. Natural History Museum of Denmark, Universitetsparken 15, Copenhagen, 2100, Denmark Author Vasilenko, Dmitry V. Cherepovets State University, Lunacharsky Prospect 5, Cherepovets, Vologda Region, 162600, Russia & Paleontological Institute, Russian Academy of Sciences, Profsoyuznaya 123, Moscow, 117647, Russia text Zootaxa 2024 2024-03-01 5418 4 328 338 http://dx.doi.org/10.11646/zootaxa.5418.4.2 journal article 10.11646/zootaxa.5418.4.2 1175-5326 10730579 7BF20349-9A30-4230-8A6B-38CF6A4C144B Lubomirus victori sp. nov. ( Figs 4–6 ) Holotype , ♀ : PIN , 964/1341, Yantarnyj , Kaliningrad Oblast , Russia , Baltic amber, late Eocene , Priabonian. Amber fragment with holotype is right parallelepiped, sized 1.8: 1.8: 0.7 mm . Paratype : ♂ , NHMD 608356 , from the same locality as the holotype . Diagnosis. Fore wing with costal, subcostal, basal, marginal, and stigmal veins sclerotized; poststigmal, radial, and postmarginal veins visible as weak traces and not forming a closed radial cell, radial vein curved ( Fig. 5B ). Mesopleuron with epicnemial carina developed only antero-ventrally ( Fig. 4E ). Description. Female ( holotype ). Body length 1.1 mm ; antennae about 0.6 times as long as body; wing 0.7 times as long as body. FIGURE 4. Lubomirus victori , holotype, female. A—whole insect in lateral view; B, E—body, lateral views; C—fore wing and metasoma, dorsal view; D—body, dorsal view. Scale bar: A—0.5 mm; B–E—0.3 mm. Head. Head smooth and bare, in dorsal view 1.9 times as wide as long, slightly wider than mesosoma; POL: 16; LOL: 9; OOL: 12; ocelli large, LOL as long as diameter of lateral ocellus; eye large and bare, 3.1 times as high as malar distance; face and cheek smooth with a few short setae; clypeus convex, transverse, 2.4 times as wide as high; epistomal sulcus distinct; tentorial pits tiny ( Fig. 5C blue arrow); mandibles elongate, narrowed to the apex, symmetrical, bidentate. Antenna. Filiform, stout; A1 cylindrical, 2.6 times as long as wide, covered with coriaceous sculpture and scattered, short setae; A3 slightly shorter than A4, A6–A8 as long as wide, A9–A14 1.12 times as long as wide, A15 2.2 times as long as wide; A3–A15 covered with short pubescence and each with several appressed thickened sensillae ( Fig. 4B ). FIGURE 5. Lubomirus victori , holotype, female. A—fore tarsus, B—fore wing; C—face; D—body, ventral view. Vein name: m —marginal, pm —postmarginal, ps —poststigmal, r —radial, s —stigmal. Scale bar 0.3 mm. FIGURE 6. Lubomirus victori , paratype, male. A—lateral view; B—dorsal view. Mesosoma. Pronotum coarsely rugose between epomia in dorsal view; pronotal shoulders angled; lateral side of pronotum smooth posteriorly from epomia and sculptured along the vertical branch of epomia; mesoscutum smooth and convex; notauli complete, crenulate; humeral sulcus longer than tegula, crenulate; scutellum convex, sculptured along posterior margin ( Fig. 4D ) (other characters of scutellum invisible); metanotum narrow, bare and coarsely rugose, metascutellum with three longitudinal keels, median keel prominent ( Fig. 4B ); propodeum in dorsal view, short, almost bare, coarsely rugose, with two elongate carinae medially ( Fig. 4D ); propleuron sculptured anteriorly, smooth and bare posteriorly; mesopleuron in lateral view with deep mesepimeral sulcus, sculpture antero-dorsally below tegula and with distinct epicnemial carina ( Fig. 4E ); mesopleuron in ventral view with acetabular carina behind fore-coxa and crenulate mesodiscrimen; metapleuron and propodeum coarsely rugose laterall with few short, scattered setae; fore tarsal claws with a row of strong setae ventrally ( Fig. 5A , red arrows), mid and hind claws simple; fore and mid legs slender; hind tibiae gradually swollen with carina on inner side. Metasoma. Petiole transverse, 1.6 times as wide as long, rugulose; sutures between tergites complete and deeply impressed; T2 convex in lateral view, entirely smooth with few scattered setae; T3–T6 short, with a row of setigerous punctures, T7 smooth, T8 with weakly prominent cerci. Base of S2 invisible, posterior part of S2 and S3–S6 smooth and almost bare, S3–S5 short, S6 subtrangular. Male. Body length 1.1 mm , antennae 0.6 as long as body. Similar to female, but antennae more slender and covered with long setae, A3 and A4 not modified, A3 slightly shorter than A2 and A4 ( Fig. 6A ), A4–A13 about 2.5 times as long as wide ( Fig. 6B ); fore tarsal claws without setae ventrally; T8 with exposed cerci. Etymology. The new species is named in honor of Victor Kolyada, enthusiastic entomologist and expert on Diaprioidea and Proctotrupoidea .