Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species
Author
Frisch, Johannes
Author
Narakusumo, Raden Pramesa
MUseUm ZoologicUm Bogoriense, Research Center for Biosystematics and EvolUtion, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor KM 46, Cibinong, West Java, 16911, Indonesia
text
Soil Organisms
2022
2022-04-01
95
1
23
73
http://dx.doi.org/10.25674/so95iss1id311
journal article
10.25674/so95iss1id311
2509-9523
10724236
Scopaeus borneensis
Cameron, 1941
(
Figs 26
,
180, 181
)
Scopaeus borneensis
Cameron, 1941: 229
.
Type specimen examined:
Holotype
♀
,
Kalimantan Barat
, Pontianak; labeled ‘Type’ (round, printed label with red margin), ‘Pontianak / Borneo’ (handwritten), ‘S. / borneensis / TYPE
Cam
.’ (handwritten), ‘M. Cameron./ Bequest./ B.M.1955-147.‘ (printed), ‘
Holotype
/
Scopaeus borneensis
/
Cameron 1941
/ label by J.
Frisch, 2022
’ (red, printed) (
NHML
).
Cameron (1941: 229)
described
Scopaeus borneensis
based on a ‘unique’ specimen. Thus, it is a
holotype
by monotypy (
ICZN
, Article 73.1.2)
.
Redescription:
Habitus and coloring as in
Fig. 26
. Head subquadrate. Penultimate antennal segment slightly elongate. Mesotibia moderately thickened. Body surface subnitid, finely, densely punctate, without microreticulation. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Forebody reddish dark brown; abdomen and antennae medium brown; maxillary palpi and legs light brown. Total body length
3.8 mm
; forebody length
2.4 mm
.
Male: Unknown.
Female: Chamber segment of sperm pump short, without apophysis, acute-angled attached to distal process with strongly widened apex (
Fig. 181
); bursal duct strong, with S-shaped portion gradually widened, then strongly narrowed towards sperm pump (
Fig. 180
).
Figures 134–145.
Abdominal sternite VII (
138
,
140
,
142
) and abdominal sternite VIII (
134 – 138
,
139
,
141
,
143– 145
) of
♂
of
Scopaeus cuspilobatus
,
holotype
,
Jawa Barat
, Sukabumi: Sirnarasa (
134
);
S. velifer
,
holotype
,
Banten
, Lebak: Citorek
Kidul
(
135
);
S. elegantulus
,
Jawa Barat
, Sukabumi, Sirnarasa: Cisarua (
136
);
S. uncinatus
,
holotype
,
Banten
, Lebak: Citorek
Kidul
(
137
);
S. bipenicillatus
,
paratype
,
Sulawesi Tengah
, Sigi: Pagana (
138
);
S. bipenicillatus
,
holotype
,
Sulawesi Tengah
, Sigi: Pagana (
139
);
S. bipectenatus
,
holotype
,
Sulawesi Tengah
, Poso: Lake Poso (
140
,
141
);
S. cuspidatus
,
holotype
,
Bali
, Jembrana: Cekik (
142
,
143
);
S. crassipunctatus
,
holotype
,
Jawa Barat
, Sukabumi, Sirnarasa: Cisarua (
144
);
S. sulawesianus
,
holotype
,
Sulawesi Selatan
: Engrekang (
145
).
Figures 146 – 154.
Sperm pump (
146 – 154
) and bursa (
148
,
149
) of ♀ of
Scopaeus javanus
, Jawa Barat, Sukabumi: Kiara Dua (
146
);
S. batukaruensis
, paratype, Bali, Tabanan: Batukaru (
147
);
S. jacobsoni
, Jawa Barat, Sukabumi: Cidahu (
148
, with lateral gonocoxal plates IX;
150
);
S. mixtus
, Malaysia, Sabah: Lokan (
149
, with lateral gonocoxal plates IX;
151
);
S. halimunsalakensis
, paratype, Jawa Barat, Sukabumi: Sirnaresmi (
152
);
S. tortuosiflagellatus
, paratype, Jawa Barat, Sukabumi: Cisalimas (
153
, with bursal duct and bursa;
154
). Long scale bar:
146
,
147
,
150 – 152
,
154
; short scale bar:
148
,
149
,
153
.
Figures 155–162.
Sperm pump (
155 – 162
) and bursa (
155
,
159
,
161
) of ♀ of
Scopaeus posoanus
, paratype, Sulawesi Tengah: Lake Poso (
155
, with lateral gonocoxal plates IX;
156
);
S
.
spiraliflagellatus
, paratype, Sulawesi Tengah: Lake Poso (
157
, with bursal duct;
158
);
S
.
anuliflagellatus
, paratype, Jawa Barat, Sukabumi: Cikaniki (
159
, with lateral gonocoxal plates folded laterally;
160
);
S
.
pulcher
, paratype, Jawa Barat, Sukabumi: Cikaniki (
161
,
162
). Long scale bar:
156
,
158
,
160
,
162
; short scale bar:
155
,
157
,
159
,
161
.
Abbreviations
:
ap
– apophysis of chamber segment,
b
– (sclerotized end of) bursa,
bd
– bursal duct,
cs
– chamber segment,
dp
– distal process of chamber segment,
lt
– lateral gonocoxal plate IX,
sd
– spermathecal duct.
Figures 163–172.
Sperm pump (
163 – 172
) and bursa (
165
,
167
,
170
,
172
) of ♀ of
Scopaeus grandis
, paratype, Jawa Barat, Sukabumi: Cidahu (
163
);
S. heronifer
, paratype, Jawa Barat, Sukabumi: Sirnarasa (
164
);
S
.
diversilobatus
, paratype, Jawa Barat, Sukabumi, Djampang Kulon (
165
, with lateral gonocoxal plates IX folded laterally;
166
);
S
.
cuspilobatus
, paratype, Jawa Barat, Sukabumi: Sirnarasa (
167
, with lateral gonocoxal plates IX,
168
);
S
.
velifer
, paratype, Banten, Lebak: Citorek Kidul (
169
);
S. elegantulus
, Jawa Barat, Sukabumi: Djampang Kulon (
170
, with lateral gonocoxal plates IX,
171
),
S. uncinatus
, paratype, Banten, Lebak: Citorek Kidul (
172
). Long scale bar:
163
,
164
,
166
,
168
,
169
,
171
,
172
; short scale bar:
165
,
167
,
170
.
Figures 173–178.
Sperm pump (
173 – 178
) and bursa (
173
) of ♀ of
Scopaeus bipenicillatus
, paratype, Sulawesi Tengah, Sigi: Pagana (
173
, with lateral gonocoxal plates IX;
174
);
S
.
bipectenatus
, paratype, Sulawesi Tengah: Lake Poso (
175
);
S. cuspidatus
, paratype, Jawa Barat: Mt Salak (
176
);
S. crassipunctatus
, paratype, Jawa Barat, Sukabumi: Sirnarasa (
177
);
S. sulawesianus
, paratype, Central Sulawesi, Tambarana (
178
, with bursal duct). Long scale bar:
174 – 178
; short scale bar:
173
.
Figures 179–181.
Sperm pump (
179
,
181
) and bursal duct (
180
) of ♀ of
S. niger
, lectotype, Singapore (
179
);
S. borneensis
, holotype, Kalimantan Barat: Pontianak (
180
, with lateral gonocoxal plates;
181
). Long scale bar:
179
,
181
; short scale bar:
180
.
Phylogeny:
Scopaeus borneensis
must be classified in the subgenus
Hyperscopaeus
Coiffait, 1984
(see
Frisch et al. 2002: 38
).
Distribution:
This species is known only from the
type
locality in
Kalimantan Barat
, Indonesian Borneo.