<document ID-DOI="10.11646/zootaxa.5263.2.4" ID-GBIF-Dataset="f6701c34-91f2-48c3-9ed1-4dd17fb3fb0d" ID-ISSN="1175-5326" ID-Zenodo-Dep="7801106" ID-ZooBank="4E687740-045F-44C5-810D-68C6EE9072C4" IM.illustrations_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" approvalRequired="4" approvalRequired_for_textStreams="4" checkinTime="1680678286118" checkinUser="plazi" docAuthor="Qi, Zhi-Hao, He, Hao, Su, Rong-Xiang, Ai, Hong-Mu &amp; Song, Hai-Tian" docDate="2023" docId="03B51633FF93FFC15FFB583C0342F83D" docLanguage="en" docName="zootaxa.5263.2.4.pdf" docOrigin="Zootaxa 5263 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5263.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lucanus ludivinae Boucher 1998" docType="treatment" docVersion="4" lastPageNumber="253" masterDocId="FF8C6E4BFF9EFFCC5F6C5E420045FFC7" masterDocTitle="Revision of the Lucanus boileaui group and notes on. takeoi Adachi, 2020 from the. maculifemoratus group (Coleoptera: Lucanidae: Lucaninae)" masterLastPageNumber="260" masterPageNumber="240" pageNumber="253" updateTime="1681158889771" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>Revision of the Lucanus boileaui group and notes on. takeoi Adachi, 2020 from the. maculifemoratus group (Coleoptera: Lucanidae: Lucaninae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Qi, Zhi-Hao</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8596-0913</mods:nameIdentifier>
<mods:affiliation>College of Plant Protection, Fujian Agriculture and Forestry University, Fuzhou, Fujian, 350002, China. &amp; Fujian Academy of Forestry, Fuzhou, Fujian, 350012, China. &amp; qizhihao 2021 @ 126. com; https: // orcid. org / 0000 - 0001 - 8596 - 0913</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>He, Hao</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-8871-9671</mods:nameIdentifier>
<mods:affiliation>College of Life Sciences, Yunnan Normal University, Kunming, Yunnan, 650500, China. &amp; 18835130675 @ 163. com; https: // orcid. org / 0000 - 0002 - 8871 - 9671</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Su, Rong-Xiang</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-6575-5776</mods:nameIdentifier>
<mods:affiliation>Fujian Academy of Forestry, Fuzhou, Fujian, 350012, China. &amp; surongxiang 2022 @ 126. com; https: // orcid. org / 0000 - 0002 - 6575 - 5776</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ai, Hong-Mu</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-7754-8256</mods:nameIdentifier>
<mods:affiliation>College of Plant Protection, Fujian Agriculture and Forestry University, Fuzhou, Fujian, 350002, China. &amp; aihongmu @ 163. com; https: // orcid. org / 0000 - 0002 - 7754 - 8256</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Song, Hai-Tian</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-1042-7959</mods:nameIdentifier>
<mods:affiliation>Fujian Academy of Forestry, Fuzhou, Fujian, 350012, China. &amp; haitiansong @ 126. com; https: // orcid. org / 0000 - 0003 - 1042 - 7959</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-04-05</mods:number>
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<mods:number>5263</mods:number>
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<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5263.2.4</mods:url>
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<subSubSection box="[151,561,1661,1688]" pageId="13" pageNumber="253" type="nomenclature">
<paragraph blockId="13.[151,689,1661,1724]" box="[151,561,1661,1688]" pageId="13" pageNumber="253">
<heading bold="true" box="[151,561,1661,1688]" fontSize="11" level="1" pageId="13" pageNumber="253" reason="1">
<taxonomicName ID-CoL="6QP72" authority="Boucher, 1998" authorityName="Boucher" authorityYear="1998" box="[151,561,1661,1688]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="253" phylum="Arthropoda" rank="species" species="ludivinae">
<emphasis bold="true" box="[151,561,1661,1688]" pageId="13" pageNumber="253">
<emphasis bold="true" box="[151,376,1661,1688]" italics="true" pageId="13" pageNumber="253">Lucanus ludivinae</emphasis>
<bibRefCitation author="Boucher, S." box="[384,561,1661,1687]" pageId="13" pageNumber="253" pagination="177 - 185" refId="ref7749" refString="Boucher, S. (1998) Deux Lucanus inedits des provinces chinoises du Yunnan et du Guangxi (Coleoptera, Lucanidae). Coleopteres, 4, 177 - 185." type="journal article" year="1998">Boucher, 1998</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,689,1697,1724]" pageId="13" pageNumber="253" type="description">
<paragraph blockId="13.[151,689,1661,1724]" box="[151,689,1697,1724]" pageId="13" pageNumber="253">
<figureCitation box="[151,284,1697,1723]" captionStart="FIGURE 1" captionStartId="2.[151,250,1840,1864]" captionTargetBox="[158,1432,666,1804]" captionTargetId="figure-216@2.[151,1436,666,1814]" captionTargetPageId="2" captionText="FIGURE 1. Male habitus of Lucanus species. A–B, L. jiaozishanus Qi, He, Su &amp; Song, new species (33.3 mm, holotype, large sized, Kunming, Yunnan); C–D, L. ludivinae Boucher, 1998 (31.3 mm, medium sized, Gongshan, Yunnan); E–F, L. takeoi Adachi, 2020 (29.2 mm, small sized, Huidong, Sichuan); G–H, L. boileaui Planet, 1897 (41.9 mm, small sized, Mabian, Sichuan). A, C, E, G, dorsal view; B, D, F, H, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7801110" httpUri="https://zenodo.org/record/7801110/files/figure.png" pageId="13" pageNumber="253">Figs. 1C–D</figureCitation>
; 
<figureCitation box="[295,364,1697,1723]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1431,181,1873]" captionTargetId="figure-17@3.[151,1435,181,1873]" captionTargetPageId="3" captionText="FIGURE 2. Male head (A, G, M, S), clypeolabrum (B, H, N, T), canthus (C, I, O, U) and tibia (D–F, J–L, P–R, V–X) of Lucanus species in dorsal view. A–F, L. jiaozishanus Qi, He, Su &amp; Song, new species (holotype, Kunming, Yunnan); G–L, L. ludivinae Boucher, 1998 (Gongshan, Yunnan); M–R, L. takeoi Adachi, 2020 (Huidong, Sichuan); S–X, L. boileaui Planet, 1897 (Mabian, Sichuan). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7814599" httpUri="https://zenodo.org/record/7814599/files/figure.png" pageId="13" pageNumber="253">2G–L</figureCitation>
; 
<figureCitation box="[375,444,1697,1723]" captionStart="FIGURE 3" captionStartId="4.[151,250,1909,1933]" captionTargetBox="[314,1276,189,1873]" captionTargetId="figure-19@4.[307,1280,181,1885]" captionTargetPageId="4" captionText="FIGURE 3. Male abdominal tergite 8 (A, E, I, M), abdominal ventrite 8 (B, F, J, N) and abdominal segment 9 (C–D, G–H, K–L, O–P) of Lucanus species. A–D, L. jiaozishanus Qi, He, Su &amp; Song, new species (holotype, Kunming, Yunnan); E–H, L. ludivinae Boucher, 1998 (Gongshan, Yunnan); I–L, L. takeoi Adachi, 2020 (Huidong, Sichuan); M–P, L. boileaui Planet, 1897 (Mabian, Sichuan). A, D, E, H, I, L, M, P, dorsal view; B–C, F–G, J–K, N–O, ventral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7814603" httpUri="https://zenodo.org/record/7814603/files/figure.png" pageId="13" pageNumber="253">3E–H</figureCitation>
; 4B, I–L; 
<figureCitation box="[557,629,1697,1723]" captionStart="FIGURE 6" captionStartId="7.[151,250,1871,1895]" captionTargetBox="[151,1433,181,1847]" captionTargetId="figure-17@7.[151,1436,181,1847]" captionTargetPageId="7" captionText="FIGURE 6. Female head (A, G, I, O), clypeolabrum (B, J, P), canthus (C, H, K, Q) and tibia (D–F, L–N, R–T) of Lucanus species in dorsal view. A–F, L. jiaozishanus Qi, He, Su &amp; Song, new species (paratype, Kunming, Yunnan); G–H (refer to p. 22 in Huang &amp; Chen, 2017; Gongshan, Yunnan); I–N, L. takeoi Adachi, 2020 (Huidong, Sichuan); O–T, L. boileaui Planet, 1897 (Mabian, Sichuan). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7814605" httpUri="https://zenodo.org/record/7814605/files/figure.png" pageId="13" pageNumber="253">6G–H</figureCitation>
; 13A
</paragraph>
</subSubSection>
<subSubSection box="[151,550,1766,1790]" pageId="13" pageNumber="253" type="reference_group">
<paragraph blockId="13.[151,550,1766,1790]" box="[151,550,1766,1790]" pageId="13" pageNumber="253">
<treatmentCitationGroup box="[151,550,1766,1790]" pageId="13" pageNumber="253">
<treatmentCitation author="Boucher, S." box="[151,547,1766,1790]" page="178" pageId="13" pageNumber="253" year="1998">
<taxonomicName ID-CoL="6QP72" authority="Boucher, 1998: 178" authorityName="Boucher" authorityPageNumber="178" authorityYear="1998" box="[151,547,1766,1790]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="253" phylum="Arthropoda" rank="species" species="ludivinae">
<emphasis box="[151,337,1766,1790]" italics="true" pageId="13" pageNumber="253">Lucanus ludivinae</emphasis>
<bibRefCitation author="Boucher, S." box="[344,547,1766,1790]" pageId="13" pageNumber="253" pagination="177 - 185" refId="ref7749" refString="Boucher, S. (1998) Deux Lucanus inedits des provinces chinoises du Yunnan et du Guangxi (Coleoptera, Lucanidae). Coleopteres, 4, 177 - 185." type="journal article" year="1998">Boucher, 1998: 178</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="253" type="materials_examined">
<paragraph blockId="13.[151,1436,1836,2043]" box="[151,1034,1836,1863]" pageId="13" pageNumber="253">
<materialsCitation box="[151,1030,1836,1863]" country="China" location="Eastern Gaoligongshan" municipality="Baoshan City" pageId="13" pageNumber="253" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis bold="true" box="[151,310,1836,1862]" pageId="13" pageNumber="253">
<typeStatus box="[151,210,1836,1862]" pageId="13" pageNumber="253">Type</typeStatus>
locality.
</emphasis>
<collectingCountry box="[316,386,1836,1862]" name="China" pageId="13" pageNumber="253">China</collectingCountry>
, 
<collectingRegion box="[396,591,1836,1863]" country="China" name="Yunnan" pageId="13" pageNumber="253">Yunnan Province</collectingRegion>
, 
<collectingMunicipality box="[602,755,1836,1862]" pageId="13" pageNumber="253">Baoshan City</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:03B51633FF93FFC15FFB583C0342F83D:8EC3F1FEFF93FFC15D92596F0443F881" box="[766,1030,1836,1863]" country="China" municipality="Baoshan City" name="Eastern Gaoligongshan" pageId="13" pageNumber="253" stateProvince="Yunnan">Eastern Gaoligongshan</location>
</materialsCitation>
.
</paragraph>
<paragraph blockId="13.[151,1436,1836,2043]" pageId="13" pageNumber="253">
<emphasis bold="true" box="[199,435,1872,1898]" pageId="13" pageNumber="253">Material examined.</emphasis>
<materialsCitation collectingDate="2015-07" collectionCode="CZHQ" collectorName="Xin-Xiong Deng" country="China" county="Nujiang" elevation="2600" location="Lushui City" municipality="Lisu Autonomous Prefecture" pageId="13" pageNumber="253" specimenCount="1" stateProvince="Yunnan">
<emphasis bold="true" box="[445,548,1872,1898]" pageId="13" pageNumber="253">
<collectingCountry box="[445,543,1872,1898]" name="China" pageId="13" pageNumber="253">CHINA</collectingCountry>
:
</emphasis>
1♁ (
<collectionCode box="[610,692,1872,1898]" pageId="13" pageNumber="253">CZHQ</collectionCode>
), 
<collectingRegion box="[713,910,1872,1899]" country="China" name="Yunnan" pageId="13" pageNumber="253">Yunnan Province</collectingRegion>
, 
<collectingCounty box="[925,1015,1872,1898]" pageId="13" pageNumber="253">Nujiang</collectingCounty>
of the 
<collectingMunicipality box="[1102,1432,1872,1898]" pageId="13" pageNumber="253">Lisu Autonomous Prefecture</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:03B51633FF93FFC15FFB583C0342F83D:8EC3F1FEFF93FFC15FFB59360158F849" box="[151,285,1908,1934]" country="China" county="Nujiang" municipality="Lisu Autonomous Prefecture" name="Lushui City" pageId="13" pageNumber="253" stateProvince="Yunnan">Lushui City</location>
, 
<location LSID="urn:lsid:plazi:treatment:03B51633FF93FFC15FFB583C0342F83D:8EC3F1FEFF93FFC15E4B59360184F848" box="[295,449,1908,1935]" country="China" county="Nujiang" municipality="Lisu Autonomous Prefecture" name="Pianma Town" pageId="13" pageNumber="253" stateProvince="Yunnan">Pianma Town</location>
<emphasis bold="true" box="[455,554,1909,1932]" pageId="13" pageNumber="253">[HḢDz</emphasis>
], 
<location LSID="urn:lsid:plazi:treatment:03B51633FF93FFC15FFB583C0342F83D:8EC3F1FEFF93FFC15D2C59360343F849" box="[576,774,1908,1934]" country="China" county="Nujiang" municipality="Lisu Autonomous Prefecture" name="Gangfang Village" pageId="13" pageNumber="253" stateProvince="Yunnan">Gangfang Village</location>
[
<emphasis bold="true" box="[789,873,1909,1932]" pageId="13" pageNumber="253">ṀĦḤ</emphasis>
], alt. 
<quantity box="[938,1024,1908,1935]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="13" pageNumber="253" unit="m" value="2600.0">
<elevation box="[938,1024,1908,1935]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="13" pageNumber="253" unit="m" value="2600.0">2600 m</elevation>
</quantity>
, 
<date box="[1035,1140,1908,1935]" pageId="13" pageNumber="253" value="2015-07">
<collectingDate box="[1035,1140,1908,1935]" pageId="13" pageNumber="253" value="2015-07">VII.2015</collectingDate>
</date>
, 
<collectorName box="[1151,1339,1908,1935]" pageId="13" pageNumber="253">Xin-Xiong Deng</collectorName>
leg.
</materialsCitation>
; 
<materialsCitation collectingDate="2018-06-30" collectionCode="CZHQ" collectorName="Ying-Bing Li" country="China" county="Gongshan County" elevation="2800" location="Lisu Autonomous Prefecture" municipality="Nujiang" pageId="13" pageNumber="253" specimenCount="1" stateProvince="Yunnan">
1♁ (
<collectionCode box="[159,241,1944,1970]" pageId="13" pageNumber="253">CZHQ</collectionCode>
), 
<collectingRegion box="[259,348,1945,1971]" country="China" name="Yunnan" pageId="13" pageNumber="253">Yunnan</collectingRegion>
, 
<collectingMunicipality box="[359,449,1944,1970]" pageId="13" pageNumber="253">Nujiang</collectingMunicipality>
of the 
<location LSID="urn:lsid:plazi:treatment:03B51633FF93FFC15FFB583C0342F83D:8EC3F1FEFF93FFC15D7D59DA0312F875" box="[529,855,1944,1970]" country="China" county="Gongshan County" municipality="Nujiang" name="Lisu Autonomous Prefecture" pageId="13" pageNumber="253" stateProvince="Yunnan">Lisu Autonomous Prefecture</location>
, 
<collectingCounty box="[866,1069,1944,1970]" pageId="13" pageNumber="253">Gongshan County</collectingCounty>
[
<emphasis bold="true" box="[1086,1170,1945,1968]" pageId="13" pageNumber="253">ỮƜB</emphasis>
], 
<location LSID="urn:lsid:plazi:treatment:03B51633FF93FFC15FFB583C0342F83D:8EC3F1FEFF93FFC15BC659DA0511F875" box="[1194,1364,1944,1970]" country="China" county="Gongshan County" municipality="Nujiang" name="Gaoligongshan" pageId="13" pageNumber="253" stateProvince="Yunnan">Gaoligongshan</location>
[
<emphasis bold="true" pageId="13" pageNumber="253">ṘN ỮƜ</emphasis>
], alt. 
<quantity box="[273,360,1980,2007]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="13" pageNumber="253" unit="m" value="2800.0">
<elevation box="[273,360,1980,2007]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="13" pageNumber="253" unit="m" value="2800.0">2800 m</elevation>
</quantity>
, 
<date box="[371,501,1980,2006]" pageId="13" pageNumber="253" value="2018-06-30">
<collectingDate box="[371,501,1980,2006]" pageId="13" pageNumber="253" value="2018-06-30">30.VI.2018</collectingDate>
</date>
, 
<collectorName box="[512,662,1980,2006]" pageId="13" pageNumber="253">Ying-Bing Li</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection box="[199,775,2016,2043]" pageId="13" pageNumber="253" type="distribution">
<paragraph blockId="13.[151,1436,1836,2043]" box="[199,775,2016,2043]" pageId="13" pageNumber="253">
<emphasis bold="true" box="[199,353,2016,2042]" pageId="13" pageNumber="253">Distribution.</emphasis>
<collectingCountry box="[359,426,2016,2042]" name="China" pageId="13" pageNumber="253">China</collectingCountry>
(
<collectingRegion box="[441,533,2016,2042]" country="China" name="Yunnan" pageId="13" pageNumber="253">Yunnan</collectingRegion>
); 
<collectingCountry box="[552,661,2017,2043]" name="Myanmar" pageId="13" pageNumber="253">Myanmar</collectingCountry>
(
<collectingRegion box="[676,763,2016,2042]" country="Myanmar" name="Kachin" pageId="13" pageNumber="253">Kachin</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>