Two new species of Begonia (section Petermannia, Begoniaceae) from the Zamboanga Peninsula, Philippines, and a redescription of Begonia parvilimba
Author
Mazo, Kean Roe F.
0000-0002-5131-9084
Forest Biological Sciences Department, College of Forestry and Environmental Science, Central Mindanao University, Bukidnon, 8714, Philippines. & keanmaze @ gmail. com; https: // orcid. org / 0000 - 0002 - 5131 - 9084
keanmaze@gmail.com
Author
Rubite, Rosario R.
0000-0002-1704-1533
Department of Biology, College of Arts and Sciences, University of the Philippines Manila, Padre Faura, Manila, Philippines & rrrubite @ up. edu. ph; https: // orcid. org / 0000 - 0002 - 1704 - 1533
rrrubite@up.edu.ph
text
Phytotaxa
2022
2022-03-07
538
2
163
171
journal article
20283
10.11646/phytotaxa.538.2.9
ab326425-fd43-4091-92a8-a63e7c40c121
1179-3163
6333742
1.
Begonia amamampang
Mazo & Rubite
,
sp. nov.
§
Petermannia
(
Fig. 2
)
Type:—
PHILIPPINES
.
Mindanao
,
Zamboanga del Norte Province
,
Municipality of Leon
B
.
Postigo
,
Brgy. Tinuyop
,
Sitio Binwangan
,
elevation
498 m
,
8°3’23.67” N
,
122°56’57.06” E
,
28 October 2021
,
K
.
R
.
F
.
Mazo
55
(
holotype
PNH
)
.
Herb
monoecious, perennial, terrestrial, up to
75 cm
tall.
Stem
branched, erect and arching,
3–6 mm
in diameter, green or reddish brown, strigose, internodes 1.0–
4.5 cm
long.
Stipules
deciduous, green, lanceolate, 9–15 × 2.5–4.0 mm, sparsely pilose, margin ciliate, apex cuspidate.
Leaves
alternate; petiole terete, 3–5 ×
1–4 mm
, green, strigose; lamina obovate, 5–12 ×
3–5 cm
, base cordate, inequilateral, margin distantly serrate, apex acuminate; 2 colour forms: one is adaxially green, minutely echinate between veins, abaxially scabrid, strigose on midrib and veins; another form is adaxially surface olivaceous, dark green, minutely echinate; abaxially surface puberulent, strigose on midrib and veins, reddish maroon; secondary veins 5–7 pairs.
Inflorescence
axillary, few-flowered, 1(–3).
Staminate flower
bracteoles persistent, lanceolate, 3.2–8.0 × 1.0–
2.5 mm
, green, sparsely puberulous, margin ciliate, apex caudate; pedicel 8.5– 11.0 mm long, white, minutely strigose; tepals 4, white suffused with pink; outer tepals ovate to suborbicular, 9–12 × 6.0–
7.5 mm
, adaxially glabrous, abaxially sparsely puberulent, apex obtuse to rounded; inner tepals linear, 5–7 × 1.0–
2.5 mm
, glabrous on both sides, apex acute or obtuse; androecium actinomorphic, stamens 18–30, filaments
0.3–0.8 mm
, fused at base; anthers obovoid, 1.0–
1.4 mm
, apex retuse.
Pistillate flower
bracteoles persistent, lanceolate, 3–11 × 1.3–2.0 mm, green, sparsely puberulous, margin ciliate, apex caudate; pedicel 2.5–5.0 mm long, white, puberulent; tepals 5, white suffused with pink; tepals lanceolate or broadly ovate to obovate, 8.0–15.0 × 6.1–8.0 mm, adaxially glabrous, abaxially sparsely puberulent to glabrous, apex obtuse to rounded; style 3, yellow, apically bifid, 4.0–
5.2 mm
long, stigma in spiral band and papillose all around; ovary trigonous-ellipsoid, green or white to pinkish, 9–12 × 4.5–8.0 mm (wings excluded), short hirsute; wings 3, equal, same color with the ovary, truncate distally,
7.5–10.2 mm
long,
5.4–7.1 mm
wide; locules 3, placenta bilamellate.
Capsule
trigonous-elliptic, 11–13 × 10.8–13.0 mm (wings included); pedicel
3–5.5 mm
long; wings 3, equal, truncate distally, rounded or subcordate proximally,
8.2–11.5 mm
long, 4.0–
5.3 mm
wide.
FIGURE 2.
Begonia amamampang
Mazo & Rubite. A.
Habit
and habitat;
B.
Portion of the stem showing strigose hairs;
C.
Leaf adaxial and abaxial surfaces showing green form;
D.
Leaf adaxial and abaxial surfaces showing olivaceous, dark green form;
E.
Staminate flower, front view;
F.
Staminate flower, side view;
G.
Pistillate flower, front view;
H
Pistillate flower showing the ovary;
I.
Cross-section of the ovary;
J.
Capsule. All from
K.R.F. Mazo 55
except (D) from K.R.F. Mazo 59.
Phenology:—
Observed flowering and fruiting between February to November.
Etymology:—
The epithet refers to the local name for
Begonia
species
in the
type
locality.
Distribution and Habitat:—
Begonia amamampang
is endemic to
Zamboanga del Norte
, Mindanao, where it is currently known only from its
type
locality. The plant grows on shaded to semi-shaded areas near creeks of secondary lowland tropical evergreen forest at
300–500 m
elevation.
Notes:—
Begonia amamampang
is similar to
B. affinis
Merr.
in having lanceolate stipules, short petioles, and number of tepals of both pistillate and staminate flowers. However, this new species differs in having stipules deciduous (vs. persistent); petioles scabrid (vs. tomentose); leaf base obtuse and equilateral (vs. laterally cordate); adaxially surface punctuate (vs. glabrous); few-flowered axillary inflorescence (vs. terminal); and trigonous-ellipsoid capsules (vs. turbinate).
Additional specimen examined:
Begonia amamampang
Mazo & Rubite.
PHILIPPINES
.
Mindanao
,
Zamboanga del Norte Province
,
Municipality of Leon
B.
Postigo
,
Brgy. Tinuyop
, near sitio
Tiwalos
,
elevation
518 m
,
8°2’54.75” N
,
122°55’50.12” E
,
12 November 2021
,
K
.
R
.F.
Mazo
59 (
PNH
)
.
Proposed Conservation Assessment:—
Begonia amamampang
is known only from the Municipality of Leon B. Postigo at two close locations, with three populations each of about one or three mature individuals at one location and another with four populations each of about one or two mature individuals. After 16 months of fieldwork with 5 or 6 times/month site visits, no additional populations were found. The localities are not currently protected under the country’s National Integrated Protected Areas System by the Department of Environment and Natural Resources. Given the ongoing threats such as nearby rubber plantation, shifting cultivation, illegal logging, and the small number of <50 mature individuals it is prudent to provisionally assess this species as Critically Endangered CR D.