The Nemourinae (Insecta, Nemouridae) of the eastern Nearctic
Author
Grubbs, Scott A.
0000-0003-2170-9716
Department of Biology and Center for Biodiversity Studies, Western Kentucky University, Bowling Green, Kentucky, 42101, USA Department of Biology and Monte L. Bean Life Science Museum, Brigham Young University, Provo, Utah, 84602, USA & scott. grubbs @ wku. edu; https: // orcid. org / 0000 - 0003 - 2170 - 9716
scott.grubbs@wku.edu
Author
Baumann, Richard W.
0000-0003-0838-4080
richard _ baumann @ byu. edu; https: // orcid. org / 0000 - 0003 - 0838 - 4080
richard_baumann@byu.edu
text
Zootaxa
2023
2023-06-19
5306
1
1
53
http://dx.doi.org/10.11646/zootaxa.5306.1.1
journal article
10.11646/zootaxa.5306.1.1
1175-5326
8054287
21AD4F85-E313-47CB-BA98-9896D2B1DC05
3.
Ostrocerca complexa
(
Claassen, 1937
)
Notched Forestfly
http://lsid.speciesfile.org/urn:lsid:
Plecoptera
.speciesfile.org:TaxonName:6144
(
Figs. 25–30
)
Nemoura complexa
Claassen, 1937:43
.
Holotype
male (Cornell University Insect Collection),
Artists Brook
,
Essex Co.
,
New York
,
USA
Nemoura complexa
:
Claassen, 1940:53
Nemoura
(
Ostrocerca
)
complexa
:
Ricker, 1952:40
Ostrocerca complexa
:
Illies, 1966:217
Ostrocerca complexa
:
Zwick, 1973:344
Nemoura
(
Ostrocerca
)
complexa
:
Hitchcock, 1974:97
Ostrocerca complexa
:
Young
et al.
, 1989:259
Distribution.
Canada
: NB, NS, ON, PQ.
USA
: CT, MA, MD, ME, NY, PA, VA, WV (
DeWalt
et al.
2022
)
Male.
Macropterous. Body length
4.1–5.8 mm
, forewing length
5.5–6.6 mm
(n = 27). Gills absent. Cerci enlarged and elongated, sclerotized, medially arcuate, and terminating in a bifurcated apex (
Figs. 25–26
). Paraprocts divided into a membranous outer lobe and a sclerotized inner lobe; inner lobe is C-shaped, apically divergent, and produced distally as a triangular projection (
Fig. 22
;
Young
et al.
1989
, their fig. 6). Epiproct elongate and complex; dorsal sclerite is bifurcated apically, open distally with dark sclerotized longitudinal bands along the midline, and membranous laterally with several spine-like projections near the base and 3–4 dark spines apically (
Figs. 26–27
); ventral sclerite is membranous apically with open medial slot, sclerotized proximally, and with distinctive fork-like process that extends through the dorsal sclerites, bearing three short prongs, one proximally and two distally (
Figs. 26–27
).
Female.
Macropterous. Body length
4.6–6.5 mm
, forewing length
6.3–7.5 mm
(n = 10). Gills absent. Cerci enlarged but unmodified (
Fig. 29
). The 8
th
sternum is moderately enlarged as a subgenital plate, extending only partially over the 9
th
sternum and distally concave; medial region is markedly sclerotized (
Figs. 29–30
). The 7
th
sternum is modified distally as a large nipple-shaped process that extends at least over ¼ of the 8
th
sternum (
Figs. 29–30
).
Larva.
Undescribed.
Comments.
Ostrocerca complexa
is an Appalachian-distributed species known from
Nova Scotia
and
New Brunswick
south to
Virginia
and
West Virginia
(
DeWalt
et al.
2022
). The females of
O. complexa
and
O. prolongata
are difficult to identify with certainty without associated males.