Taxonomic review of Coccobius species (Hymenoptera: Aphelinidae) from China, with notes on their use in biological control of scale insect pests (Hemiptera: Diaspididae), and description of a new species
Author
Wang, Zhu Hong
Key Laboratory of Integrated Pest Management for Fujian-Taiwan Crops, Ministry of Agriculture, China; & Fujian Provincial Key Laboratory of Insect Ecology, China; & Fujian Agriculture and Forestry University, Fuzhou, China
Author
Huang, Jian
Key Laboratory of Integrated Pest Management for Fujian-Taiwan Crops, Ministry of Agriculture, China; & Fujian Provincial Key Laboratory of Insect Ecology, China; & Fujian Agriculture and Forestry University, Fuzhou, China
Author
Pan, Dong Ming
Fujian Agriculture and Forestry University, Fuzhou, China
text
Journal of Natural History
2013
J. Nat. Hist.
2013-08-13
48
5 - 6
359
373
http://dx.doi.org/10.1080/00222933.2013.791954
journal article
7040
10.1080/00222933.2013.791954
13137cdc-83ca-408b-bf82-995abdb2cf88
1464-5262
4632155
Genus
Coccobius
Ratzeburg, 1852
Coccobius
Ratzeburg, 1852
. Ichneum. Forstin., 3: 195.
Type
species:
Coccobius annulicornis
Ratzeburg
, by subsequent designation (
Gahan & Fagan 1923
).
Physcus
Howard, 1895
. Tech. Ser.
U.S.
Dept. Agr. Div. Ent., 1: 43.
Type
species:
Coccophagus varicornis
Howard
, by monotypy.
Encyrtophyscus
Blanchard, in
De Santis, 1948
. Rev. Mus. La Plata (N.S.), 5 Sec. Zool.: 192.
Type
species:
Physcus flavoflagellatus
De Santis
, by monotypy and original designation. Synonymized by
Hayat (1983)
.
Physculus
Jasnosh, 1977
. Zool. Zh., 56: 1115.
Type
species:
Physculus danzigae
Jasnosh
, by monotypy and original designation. Synonymized by
Hayat (1983)
.
Coccobius
species can be distinguished from all other genera of
Aphelinidae
as follows.
Female
. Body yellow to black; antenna often with some segments pale and others brown to dark brown; wings usually hyaline, rarely slightly and evenly infuscated. Antenna seven-segmented, with funicle three-segmented, clava two-segmented; mesopleuron large, undivided; mid lobe of mesoscutum large, with numerous setae, axillae small and widely separated, each with a single seta, scutellum with four to six setae. Fore wing without linea calva, often with narrow area basally asetose. Postmarginal vein absent or very short. Tarsal formula 5-5-5.
Male.
Differing from the female mainly in genitalia and antennal shape. Antenna eight-segmented, funicle not differentiated from clava, flagellum uniformly coloured.
Key to the species of
Coccobius
(females) from
China
1. Mesosoma (“thorax”) yellow to yellow-brown............................ 2 Mesosoma dark brown to black......................................... 6
2. Antenna with contrastingly coloured flagellomeres....................... 3 Antennal flagellum uniformly dark.......................
C. furviflagellatus
3. F2 dark brown.......................................................... 4 F2 pale................................................................. 5
4. F1–F3 each more than twice as long as wide....................
C. flaviceps
F1–F3 each less than twice as long as wide........................
C. fulvus
5. Scutellum with eight setae..................................
C. annulicornis
Scutellum
with four setae...............................
C. curtifuniculatus
6. Metasoma brown....................................................... 7 Metasoma pale, or sides of metasoma with dark stripes or markings laterally....................................................................... 11
7. Antennal flagellum uniformly either pale, or dark brown except clava distally pale.................................................................... 8 Antenna flagellum differently coloured................................... 9
8. Scutellum with two pairs of setae..............................
C. languidus
Scutellum
with three pairs of setae...........................
C. longialatus
9. Pedicel brown......................................................... 10 Pedicel pale....................................................
C. azumai
10. Clava pale.............................................
C. longifuniculatus
Clava dark......................................................
C. furvus
11. Metasoma uniformly pale.............................................. 12 Metasoma with dark stripes or markings laterally....................... 13
12. Pedicel pale; mid lobe of mesoscutum with 12–18 fine setae (excluding pair of robust setae posteriorly).....................................
C. abdominis
Pedicel dark; mid lobe of mesoscutum with approximately 36 fine setae (excluding pair of robust setae posteriorly.........................
C. chaoi
13. Metasoma with separated dark markings laterally....................... 14 Metasoma with continuous dark stripes laterally........................ 15
14. F1 and clava dark brown.....................................
C. maculatus
F1 and clava pale.............................................
C. wuyiensis
15. F1 and clava dark...............
C
.
albiscutellum
Wang and Huang
,
sp.nov.
F1 and clava pale............................................
C. flavicornis
Species accounts