A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae)
Author
Platnick, Norman I.
Author
Ubick, Darrell
text
American Museum Novitates
2008
2008-04-09
3608
1
1
36
http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2008)3608%5B1%3AAROTEC%5D2.0.CO%3B2
journal article
10.1206/0003-0082(2008)3608[1:AROTEC]2.0.CO;2
0003-0082
5389369
Titiotus marin
,
new species
Figures 77–81
; map 3
TYPE
:
Male
holotype
from
Mill Valley
,
Marin Co.
,
California
(
June 1951
;
D. Hardy
), deposited in
AMNH
.
ETYMOLOGY: The specific name is a noun in apposition taken from the
type
locality.
DIAGNOSIS: Males resemble those of
T. humboldt
and
T. costa
but can be distinguished by having the third most ventral prong of the tibial apophysis notched (fig. 79); females have a relatively narrow, ovoid epigynal septum (fig. 80).
MALE: Total length 11. Coloration as in
T. californicus
. Leg spination: femora: I p0-3-1, r1-2-1; II r0-2-1; III r1-1-2; IV r0-1-2; tibiae: I, II d1-0-1, v4-4-4, r1-1-0; III r0-1-1; metatarsi: I, II p1-1-0, r1-1-0; III p1-2-2; IV r1-2-1. Embolus prolaterally expanded near tip, distally narrow; median apophysis with only tiny subdistal projection; most ventral prong of tibial apophysis triangular, middle prong relatively short, distally incised, dorsal prong distally narrowed (figs. 77–79).
FEMALE: Total length 11. Coloration as in
T. californicus
. Leg spination: femora: I–III r1-1-2; IV p1-2-1, r0-1-2; tibiae: I p1-0-0, v4-4- 4, r1-0-0; II d1-0-0, p0-1-1, v4-4-4, r1-1-0; III r0-1-1; metatarsi III p1-2-2. Epigynal septum relatively narrow, ovoid, posterolateral epigynal margins protuberant; posterior ducts oblique (figs. 80, 81).
Figs. 77–81.
Titiotus marin
,
new species
.
77.
Left male palp, prolateral view.
78.
Same, ventral view.
79.
Same, retrolateral view.
80.
Epigynum, ventral view.
81.
Same, dorsal view.
OTHER MATERIAL EXAMINED:
United States
:
CALIFORNIA
:
Marin Co.
:
Burdell Mountain
, SE
slope,
Novato
, 38
°
07
9
48
0
N
, 122
°
34
9
09
0
W
,
Mar. 16, 1990
, serpentine grassland with adjacent mixed woodland (D.
Figs. 82–86.
Titiotus costa
,
new species
.
82.
Left male palp, prolateral view.
83.
Same, ventral view.
84.
Same, retrolateral view.
85.
Epigynum, ventral view.
86.
Same, dorsal view.
Ubick, T. Briggs, CDU), 13,
Mar. 8, 1991
, oak woodland, under serpentine (D. Ubick, CDU),
1♀
;
Fairfax
,
Nov. 15, 1974
(
CAS
)
,
13, 38
°
00.0
9
N, 122
°
36.0
9
W,
May 6, 1999
, in house in oak woodland (J.
Schweikert
,
CAS
)
,
13,
Mt. View Road
,
Apr. 28, 2004
(D.
Giampietro
,
CAS
)
,
13;
Marinero Estates
,
Tiburon Peninsula
,
Apr. 9, 1994
, serpentine grassland (
D. Ubick
,
CDU
)
,
23;
Mill Valley
, no date, running on floor in house (
E. Ross
,
CAS
)
,
1♀
,
June 1951
(D.
Hardy
,
AMNH
)
,
13,
June 7, 1954
(H.
Leech
,
AMNH
)
,
13,
June 17, 1955
, at light (H.
Leech
,
AMNH
)
,
13,
June 1956
(E.
Ross
,
AMNH
)
,
13,
Aug. 1956
, in house (E.
Ross
,
AMNH
)
,
1♀
;
Apr. 1957
, in empty bathtub at night (E.
Ross
,
AMNH
)
,
13,
June 14, 1957
, in house (H.
Leech
,
AMNH
)
,
13,
June 1, 1958
(F.
Leech
,
CAS
)
,
13,
Sept. 22, 1958
(H.
Leech
,
CAS
)
,
13,
Oct. 1, 1958
, in house (E.
Ross
,
AMNH
)
,
1♀
,
Aug. 1, 1963
, in house at night (E.
Ross
,
CAS
)
,
1♀
,
May 1965
, house basement (E.
Ross
,
CAS
)
, 23,
1♀
,
June 6, 1988
, in basement of house (C.
Hausner
,
CDU
)
,
1♀
,
Sept. 5, 1991
(L.
Alderson
,
CDU
)
,
1♀
,
June 9, 1995
(
V
.
Roth
,
CAS
), 13,
June 11, 1996
, in house (E.
Ross
,
CAS
)
,
13,
Aug. 4, 2002
(S.
Hall
,
UCR
)
,
1♀
,
May 30, 2003
(E.
May, UCR
),
1♀
;
Mount Tamalpais
,
Mar. 14, 1909
(E.
VanDyke
,
CAS
)
,
13;
Novato
,
May 6, 1973
(
B. Kessel
,
CAS
)
,
13,
Mar. 14, 1982
, under serpentine (D.
Ubick
,
CDU
)
,
13,
Apr. 25, 1982
, under serpentine float (D.
Ubick
,
CDU
)
,
13,
Jan. 1990
, under serpentine (D.
Ubick
,
CDU
)
,
13;
Ring Mountain
,
Tiburon Peninsula
,
Apr. 15, 1994
, serpentine grassland (
D. Ubick
,
J. Boutin
,
CDU
)
,
13 (penultimate, but palp details visible through cuticle);
San Anselmo
,
May 22, 1998
, in house (
CAS
)
,
1♀
,
Aug. 29, 2000
, in house (D.
Burns
,
CAS
)
,
1♀
;
San Geronimo
,
Jan. 15, 1994
(matured
Mar. 24, 1994
), serpentine grassland (
D. Ubick
,
J. Boutin
,
CDU
)
,
13;
San Rafael
,
June 28, 1993
, in house (
N. Durighello
,
CDU
)
,
1♀
, mid-May 2000, in house (D.
Halliday
,
UCR
)
,
13;
N San Rafael
,
Feb. 6, 1988
(3 matured
Mar. 21, 1988
,
♀
Apr. 4, 1988
), serpentine grassland (D.
Ubick
,
CDU
)
, 13,
1♀
;
San Rafael Ridge
,
San Anselmo
,
May 27, 1996
(
L. Freihofer
,
CAS
)
,
1♀
;
Sausalito
(hilly part),
May 23, 2000
, in house (
A. Weisskopf
,
CAS
)
,
13;
South Tiburon Ridge
, 37
°
53
9
05
0
N
, 122
°
27
9
06
0
W
,
Feb. 27, 1993
, serpentine grassland (D.
Ubick
,
J. Boutin
,
C. Kellner
,
CDU
)
,
2♀
,
Apr. 18, 1993
, serpentine grassland (D.
Ubick
,
CDU
)
, 23,
1♀
;
Terra Linda
,
San Rafael
, 38
°
01.2
9
N, 122
°
34.5
9
W, all border of suburb and open space, grassy hills, oaks, laurels, shrubs, elev.
100 m
(
J. Schweikert
,
CAS
)
:
June 4, 2003
, 13,
June 22, 2003
, 13,
July 15, 2003
, 13,
July 31, 2003
,
1♀
,
Aug. 23, 2003
,
1♀
,
Apr. 13, 2004
, 13,
Apr. 25, 2004
, 13,
May 2, 2004
, 13,
July 18, 2004
, 13,
May 29, 2005
, 13,
May 13, 2006
, 13;
Tiburon
,
Dec. 21, 1988
(L.
Cheng
,
T
.
Briggs
,
CAS
)
,
1♀
, mid-April 2002 (K.
Passaretti
,
UCR
)
,
23;
Woodacre
,
Apr. 21, 1981
, in house (
P. Signet
,
UCB
)
,
13.
Mendocino Co.
:
Ukiah
,
June 5, 2000
, in house (
E. Nicolait
,
CAS
)
,
13.
Sonoma Co.
:
Petaluma
,
May 16, 2003
, in bathroom (
H. Stavropoulos
,
UCR
)
, 13.
DISTRIBUTION: Coastal
California
, from the
San Francisco Bay area
north (map 3)
.