On the type selection and re-typification of two monitor lizard taxa (Squamata: Varanidae): Monitor bivittatus celebensis Schlegel, 1844 and Monitor kordensis Meyer, 1874; with some comments and corrections on other name-bearing type specimens
Author
Böhme, Wolfgang
Author
Koch, André
text
Zootaxa
2010
2440
60
68
journal article
10.5281/zenodo.294103
fb1c6659-538b-4a8b-821f-44d50b91af9f
1175-5326
294103
1.
Monitor bivittatus celebensis
Schlegel, 1844
This taxon described from Northern Celebes (= Sulawesi) was meanwhile considered to be a synonym of
Varanus salvator salvator
(Laurenti, 1768)
(see
Mertens 1942
,
1963
;
Böhme 1997
,
2003
) implying that the likewise Sulawesian taxon
Monitor
(
Hydrosaurus
)
togianus
Peters, 1872
(at present
V. togianus
) would be restricted to its
type
locality, the off-coast Togian Islands of Central Sulawesi. But already since the end of the 19th century, there were hints that
togianus
-like, melanistic phenotypes would occur also on Sulawesi itself (
Weber 1891
;
Boulenger 1897
;
Iskandar & Nio 1996
;
Auliya 2006
;
Koch
et al.
2007
) implying the sympatric existence of melanistic (
togianus
) and patterned (
celebensis
) phenotypes, thus, in case of two distinct taxa, even the coexistence of two distinct species. Under these auspices,
De Lisle (2009)
felt obliged ("it seems wise to recognize a
type
.....") to identify a
lectotype
from, as he thought, only two RMNH specimens from Sulawesi because
Schlegel (
1837-44
)
did not fix a name-bearing
type
, neither by original designation nor by implication. According to Art. 74 (ICZN 1999) this action is possible although unnecessary in a checklist without a revisionary taxonomic approach which, however, is in progress by one of us (AK). Of the supposed two historical RMNH specimens from Sulawesi available to Herrmann Schlegel during the time of his curatorship at the Leiden museum,
De Lisle (2009)
selected the specimen RMNH 3179 (an adult, collected by E. A. Forsten, 1841) because it had a specific locality (“Menado” = Manado, North Sulawesi) while the second (RMNH 3176, collected by v[an]. Delden, without date) was labelled just as "Celebes". Here, it should be mentioned that collection number RMNH 3176 represents not one female as
De Lisle (2009)
stated, but actually comprises two juvenile specimens (see
Brandenburg 1983
: 59;
Koch
et al.
2007
: 152). De Lisle’s (2009) conclusion that specimens RMNH 3176a, b were already present in the Leiden museum in the early 1840s, probably due to the low accession number, is also incorrect because the RMNH collection numbers were not allocated chronologically (E. Gassó-Miracle, pers. comm.). According to a note in the catalogue by the former curator of herpetology at the Leiden Museum, Rinus Hoogmoed, “numbers up to ca. RMNH 3760 are classified systematically, higher numbers irregular. Up to that it concerns animals received up to ca. 1866. From RMNH 3760 irregular with older specimens (1837) and newer (1872, 1877). I think that from the end of the 1860's, beginning 1870's (RMNH 3881 and further) it seems that specimens were classified on receiving date” (R. de Ruiter, pers. comm.). The date of collecting for RMNH 3176a, b was given as “1872?” by
Koch
et al.
(2007)
. Van
Steenis-Kruseman (1950)
, in her compendium about Malaysian (plant) collectors, did mention a person “van Delden”, however, without providing any dates of birth and death or collecting trips. In sum, it is very likely that RMNH 3179 was the only Sulawesian specimen available to Schlegel for the description of his
Monitor bivittatus celebensis
,
and therefore represents the
holotype
of this taxon by implication.
Consequently, De Lisle's (2009)
lectotype
designation becomes inappropriate. He merely mentioned the unknown
holotype
specimen as had earlier been indicated by
Koch
et al.
(2007)
. If designating a
lectotype
for Schlegel's taxon was warranted,
De Lisle (2009)
correctly followed Recommendation 74B to select a specimen figured before in the literature (see Fig.
13 in
Koch
et al.
2007
) but he did not mention this previously published figure. Moreover, he ignored Recommendation 74C, because he did not publish the data described in Recommendation 73C ("Data on the
holotype
") which are likewise requested for
lectotype
designations.
Due to the minor quality of the photograph of Schlegel’s
type
of
V. s. celebensis
in
De
Lisle (2009
: 10, which shows only half of the specimen’s body) and because the publication by
Koch
et al.
(2007)
may be difficult to access, we here provide a better photograph of the
holotype
(see
Fig. 1
).